Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
In my Prodromus (1881, p. 446) I had enumerated the Semantida with three
genera (Nos. 298 to 300) as a separate subfamily of the Dyostephida or
"Stephoidea biannularia," and characterised these "Dyostephanida" by the
following definition: "Skeleto annulis duobus composito, qui in duobus
planis invicem perpendicularibus jacent; altero annulo (sagittali)
verticali, altero (basali) horizontali." As the names there given were
already employed with another signification, and as the Zygostephanida
(there united with the Dyostephanida) are more closely related to the
Coronida, I now change the names, and propose to call the family Semantida,
expressing by this term the typical similarity of the skeleton to a
signet-ring (_Semantis_, _Semantrum_, _Semantidium_).
At about the same time, some #Stephoidea# of this family were accurately
described by Bütschli (1882, Zeitschr. f. wiss. Zool., vol. xxxvi. p. 495,
Taf. xxxii. figs. 6, 7, 8). He called them _Stephanolithis_, a name which
Ehrenberg had employed, not for complete shells of Radiolaria, but for
isolated parts of such, and for siliceous fragments of different skeletons,
needles of Sponges, &c. The three species described by Bütschli represent
three different genera of our Semantida, viz., _Semantis spinescens_ (with
two gates in the basal plate), _Semantrum mülleri_ (with four gates), and
_Semantidium haeckelii_ (with six gates). He pointed out the great
morphological value of the fenestrated basal plate and its paired gates, as
beginnings of numerous other NASSELLARIA. But his opinion, that in all
#Spyroidea# and #Cyrtoidea#, derived from these, two pairs of basal gates
were constant, was erroneous, nor was the formation of the first pair
naturally explained; he supposed that the formation of the basal plate
begins by development of an odd sagittal apophysis, arising from the base
of the primary sagittal ring. But this odd sagittal apophysis ("der unpaare
mediane Kieselfortsatz c1", _loc. cit._, p. 497) is in reality not a
primary {954}and essential part of the skeleton, but secondary and of
little morphological value, absent in the majority of the Semantida and of
the other NASSELLARIA.
We divide our family Semantida into two different subfamilies, which
possibly possess a direct phylogenetic relation to the two subfamilies of
Stephanida:--The Semantiscida have no typical feet, and have arisen
directly from the Lithocircida; the Cortiniscida, however, possess the
three typical basal feet of _Cortina_, and may therefore be derived
directly from the Cortinida. Since these three cortinar feet are probably
identical with the three primary radial rods of the #Plectoidea#, an
immediate affinity also to these NASSELLARIA is indicated.
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