Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
The Semantiscida, which do not possess these three basal cortinar feet, are
the simpler forms of the family. The simplest of all, and perhaps the
common ancestral form of the whole family, is _Semantis_ (Pl. 92, figs. 1,
2). It may be derived from _Archicircus_ or _Zygocircus_ by development of
two pairs of horizontal apophyses on its base, around the porochora of the
central capsule. The two rods of each side (right and left), becoming
curved one towards the other, and meeting laterally, form a simple
horizontal gate, and the two paired basal gates together, a horizontal ring
or basal ring, to which the primary sagittal ring is perpendicular. In the
next allied genus, _Semantrum_ (Pl. 92, figs. 3-5), three pairs of
horizontal apophyses are developed, and therefore two pairs of basal gates
produced, an anterior and a posterior. In the third genus, _Semantidium_
(Pl. 92, figs. 6, 7), three pairs of basal pores or gates are visible,
surrounded and separated by four pairs of horizontal apophyses, which arise
from the base of the sagittal ring. Finally, in _Clathrocircus_ (Pl. 92,
figs. 8-10) the number of apophyses is much increased, and two parallel
rows of pores are developed along the two sides of the sagittal ring.
The basal plate or the "seal," developed from the base of the primary
sagittal ring or "signet-ring," is therefore a horizontal ring, which
becomes bisected by the latter, and exhibits either one pair of primary
"basal gates" or two or three pairs of these important basal pores, rarely
more. Since these pores possess the greatest morphological value, and are
probably everywhere homologous, we give to them and to the separating
apophyses certain names, and call the anterior pair of gates, "jugular
pores" (_i_ in our figures, the pair I of Bütschli); the middle (usually
the largest) pair, "cardinal pores" (_k_ in our figures, the pair II of
Bütschli), and the posterior, smaller pair, "cervical pores," _l_. The
typical pairs of rods, by the union of which these basal pores arise, are
the following:--(1) the clavicular or furcular rods, _f_, the first pair
(rods _e_1 of Bütschli), (2) the coracal rods, _e_, between the jugular and
cardinal pores (rods _e_ of Bütschli), (3) the scapular rods, _g_, between
the cardinal and cervical pores (rods _e_2 of Bütschli), (4) the cervical
rods, the fourth pair of apophyses, the most posterior, _h_. Bütschli
supposes that the topographical succession of the three typical pairs of
basal pores is also the chronological succession, the jugular being formed
first, the {955}cardinal second and the cervical pores third but it seems
that this succession is often altered and that the cardinal pores (the
largest), appear first, the jugular pores (in front of them) second and the
cervical pores third (or perhaps sometimes in the inverse succession).
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