Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIIIHaeckel, Ernst
Science
Report on the Radiolaria Collected by H.M.S. Challenger During the Years 1873-1876, Second Part: Subclass Osculosa; Index: Report on the Scientific Results of the Voyage of H.M.S. Challenger During the Years 1873-76, Vol. XVIII
Haeckel, Ernst
Radiolaria
C. The skeleton is composed of a latticed cephalis only, a simple ovate or
subspherical fenestrated shell, which encloses the monopylean central
capsule; there is no {893}trace of a sagittal ring nor of a basal tripod.
This is the case in the remarkable family of Cyrtocalpida (Archicorida and
Archicapsida), in numerous #Botryodea# and in other #Cyrtellaria#.
D. The skeleton is composed of a sagittal ring and a basal tripod, without
latticed cephalis. This is the case in a few, but very important forms of
#Stephoidea#: _Cortina_, _Stephanium_, _Cortiniscus_, _Stephaniscus_,
_Podocoronis_, and some allied genera.
E. The skeleton is composed of a sagittal ring and a latticed cephalis, but
without basal tripod. This is the case in numerous #Cyrtellaria#, in the
Circospyrida (or Zygospyrida apoda: _Dictyospyris_, _Circospyris_) and some
other #Spyroidea#; and in a large number of #Botryodea# and Cyrtoidea
eradiata (a part of the Sethocyrtida, Theocyrtida, Lithocampida, and
others).
F. The shell is composed of a basal tripod and a latticed cephalis, but
without any trace of the sagittal ring. This is the case in numerous
Cyrtoidea triradiata and multiradiata, and perhaps in the majority of the
following families--Tripocalpida, Tripocyrtida, Podocyrtida, and
Podocampida.
G. The shell is composed of all three above-mentioned elements, of a
sagittal ring, a basal tripod, and a latticed cephalis. This is the case in
the great majority of #Spyroidea# (with a few exceptions only), and perhaps
also in the majority of #Cyrtoidea#.
The survey of these seven groups, A to G, each of which is represented by
numerous living forms, shows clearly how difficult and complicated the
morphology and phylogeny of the numerous NASSELLARIA must be. For all
possible combinations of the three original structural elements are
realised abundantly, and in such complicated relations, and so intermingled
in the different orders and families, that it seems nearly hopeless to
answer the question of their true origin. The identity in the structure of
the central capsule, however, in all these MONOPYLEA, makes it probable
that they have all arisen originally from the skeletonless Nassellida
(_Cystidium_, _Nassella_), either in a monophyletic or in a polyphyletic
way. In this respect the following phylogenetical hypotheses are possible.
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