Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
=5. Plant succession.= The fact that formations arise and disappear was
perceived by Biberg as early as 1749, but it received slight attention
until Steenstrup’s study of the succession in the forests of Zealand
(1844 prox.). In the development of formations, as well as in their
recognition, nearly all workers have confined themselves to the
investigation of particular changes. Berg (1844), Vaupell (1851),
Hoffmann (1856), Middendorff (1864), Hult (1881), Senft (1888), Warming
(1890), and others have added much to our detailed knowledge of
formational development. Notwithstanding the lapse of more than a half
century, the study of plant successions is by no means a general
practice among ecologists. This is a ready explanation of the fact that
the vast field has so far yielded but few generalizations. Warming
(1895) was the first to compile the few general principles of
development clearly indicated up to this time. The first critical
attempt to systematize the investigation of succession was made by
Clements (1904), though this can be considered as little more than a
beginning on account of the small number of successions so far studied.
Future progress in this field will be conditioned not only by the more
frequent study of developmental problems by working ecologists, but
also, and most especially, by the application of known principles of
succession, and by the working out of new ones.
=6. Ecological phytogeography.= Until recent years, the almost universal
tendency was to give attention to formations from the standpoint of
vegetation alone. While the habitat was touched here and there by
isolated workers, and plant functions were being studied intensively by
physiologists, both were practically ignored by ecologists as a class.
The appearance of Warming’s _Lehrbuch der oecologischen
Pflanzengeographie_ (1896) and of Schimper’s _Pflanzengeographie auf
physiologischer Grundlage_ (1898) remedied this condition in a measure
by a general discussion of the habitat, and by emphasizing the
importance of the ecological or physiological point of view. Despite
their frank recognition of the unique value of the habitat, the major
part of both books was necessarily given to what may be termed the
general description of formations. For this reason, and for others
arising out of an almost complete dearth of methods of investigation,
ecology is still almost entirely a floristic study in practice, although
there is a universal recognition of the much greater value of the
viewpoint which rests upon the relation between the formation and its
habitat.
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