Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
=200. Possible objections.= The use of the quadrat has led to the
criticism that it is needlessly detailed and thorough, and that, after
all, the space covered is but a minute part of the entire formation. The
first objection is one that has also been urged against the use of
instruments of precision in the habitat. It is always brought forward by
those who have not used instruments, and as witnesses they are of
necessity incompetent. No one who is familiar with the instrument or the
quadrat by actual practice has felt that the methods based upon them
were too thorough. In no case has the writer ever listed or mapped a
quadrat without discovering some new fact or relation, or clearing up an
old question. It can not be denied that quadrat methods require both
time and patience, but this is true of any kind of research work that is
at all worth while. Every ecologist, moreover, that has the interests of
his field at heart and deprecates the present slipshod work, will
appreciate the necessity of methods which seem like drudgery to the mere
dabbler.
The second objection, that the quadrat is at best but a small bit of the
area under investigation, seems at first to be a valid one. It can not
be gainsaid that the actual space studied is insignificant as compared
with the whole formation; still, it must be obvious that even a single
quadrat can add at least some facts of value, which can never be
obtained by the best of general methods. Furthermore, if the formation
be an actual and not an imaginary one, a single quadrat will be in some
measure representative. In the more homogeneous ones, it will have much
the same value that a type specimen bears to the species established
upon it. In formations which are less uniform, its value is
correspondingly reduced, so that in formations which show marked zones,
consocies, or patches, it becomes necessary to locate a quadrat in each.
In the matter of representation alone, the graphic method of the quadrat
map with its close-focus detail photograph, is far superior to anything
that can be obtained by the ordinary description and photograph.
Finally, the scientific study and recording of succession, and
particularly of competition, is an impossibility without the aid of the
permanent and denuded quadrat. The stoutest champion of the practice of
walking through a formation, and jotting down impressions, can not avoid
their use if he would attack these problems, and, once familiar with the
quadrat, his objections to the drudgery of thoroughness will soon
vanish.
_Kinds of Quadrats and Their Use_
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