Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
=201. Size and kinds.= The unit size of quadrat is the meter, and when
the term is used without qualification, it refers to the meter quadrat.
To make them strictly comparable, and exactly divisible, unit quadrats
are always grouped in squares; thus a major quadrat is a square of four
units, and a perquadrat one of sixteen units, or four meters square.
Quadrats of greater size are necessary in woodland and forest, where the
rule, however, is that the woody plants alone are recorded for the whole
quadrat, the herbaceous growth being listed or mapped for but one or two
representative units. For special purposes, quadrats of 3, 5, 6, etc.,
meters may be used, but they are much less convenient. Quadrats are
further distinguished with respect to their use. A list quadrat is one
in which the plants are merely listed and the number of individuals of
each species indicated. Chart quadrats are those in which the area
concerned is accurately mapped on plotting paper. Both list and chart
quadrats are rendered permanent by careful labeling, so that their
changes can be followed from year to year. The greater value of the
chart causes practically all permanent quadrats to be of this type, and
for the same reason only permanent chart quadrats are converted into
denuded ones.
=202. Tapes and stakes.= The lines for marking out quadrats are made of
strong white tape, ⅝ inches wide. This is doubled and sewed firmly at
both edges. Under moderate stretching, the tape is carefully marked off
into decimeters, and eyelets 5 mm. in diameter are set in at each end
and at the marks. This can readily be done by any shoemaker at slight
expense. The usual lengths are one and two meters, as these are most
frequently used, and they can also be easily combined to make larger
quadrats. The tapes are slightly longer than one meter in order that the
distance between the end eyelets may be exact. The tapes of the larger
forest quadrats should be divided into lengths of one meter, as these
permit ready plotting and also make it possible to interpolate a meter
quadrat for the study of the undergrowth at any point. The intervals of
the tape are numbered from left to right, as conspicuously and clearly
as possible. For this a waterproof ink or paint is very desirable. For
holding the tapes in position, hatpins, nails, and meat-skewers have
been used with more or less satisfaction. The ideal stake, however, is
one which holds the tape close to the ground, and can be readily moved.
It is merely a stout wire, 3 mm. in diameter and 8 inches long, looped
at the top, sharpened at the tip, and with a small ring of solder 3
inches from the tip.
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