Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
=260.= By invasion is understood the movement of plants from an area of
a certain character into one of a different character, and their
colonization in the latter. This movement may concern an individual, a
species, or a group of species. From the nature of invasion, which
contains the double idea of going into and taking possession of, it
usually operates between contiguous formations, but it also takes place
between formational zones and patches. More rarely and less noticeably,
there may be invasion into a remote vegetation, as a result of long
carriage by wind, water, birds, railroads, or vessels. Movement or
migration, however, represents but one of the two ideas involved in
invasion. Migration merely carries the spore, seed, or propagule into
the area to be invaded. In ecesis, the spores or seeds germinate and
grow, after more or less adjustment, and in case the latter becomes
sufficiently complete, the new plants reproduce and finally become
established. With all terrestrial plants, invasion is possible only when
migration is followed by ecesis, because of the inherent differences of
formations or of areas of the same formation. In the case of surface
floating forms, such as _Lemnaceae_, and of the plancton, ecesis is of
much less importance, on account of the uniformity of the medium and the
lack of attachment, and migration is often practically synonymous with
invasion.
_MIGRATION_
=261.= Migration has been sometimes used loosely as a synonym for
invasion, but it is here employed in its proper sense of removal or
departure, i. e., movement, and is contrasted with ecesis, the making of
a home, the two ideas being combined in invasion, which is a moving into
and a taking possession of. An analysis of migration reveals the
presence of four factors, mobility, agency, proximity, and topography.
Not all of these are present in every instance of migration, as for
example in the simple elongation of a rootstalk, but in the great
majority of cases each plays its proper part. Mobility represents the
inherent capacity of a plant for migration, and in its highest
expression, motility, is in itself productive of movement. As a general
rule, however, modifications for securing mobility are ineffective in
the absence of proper agents, and the effective operation of the two
will be profoundly influenced by distance and topography.
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