Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
=266. Seed production.= The relation of spore or seed-production to
mobility is obvious in the case of mobile species; in the case of
immobile ones, it is just as evident that it has no effect, though it
may still have considerable influence in increasing migration. In the
case of two species with equally effective dissemination contrivances,
the one with the largest seed-production will be the more mobile. On the
basis of the relation of seeds to flower, two groups of plants may be
distinguished, one, _Polyanthae_, in which the flowers are many and the
seeds few or single, as in _Compositae_, and the other, _Polyspermatae_,
_Portulaca_, _Yucca_, etc., in which the number of seeds to each flower
is large. So far as the actual number of seeds produced is concerned,
polyanthous plants may not differ from polyspermatous ones, but, as a
rule, they are much more highly specialized for dissemination and are
more mobile. The number of fertile seeds is also much greater, a fact
which is of great importance in ecesis, and which, taken in connection
with mobility, partially explains the supremacy of the composites. Among
the fungi and algae, the amount of spore-production in a large degree
determines the mobility, since these forms are intrinsically permobile.
=267. Agents of migration.= In the last analysis, however, the
possibility of migration depends upon the action of distributive agents;
in the absence of these, even the most perfect contrivance is valueless,
while their presence brings about the distribution of the most immobile
form. In short, migration depends much more upon such agents than upon
mobility, however perfect the latter may be. It is, moreover, evident
that the amount and extent of migration will be determined primarily by
the permanence and forcefulness of the agent, as indicated by its
ability to bring about transportation. Finally, as will be shown later,
the direction and rapidity of migration depend directly upon the
direction and intensity of the agent.
Migration results when spores, seeds, fruits, offshoots, or plants are
moved out of their home by water, wind, animals, man, gravity, glaciers,
growth, or mechanical propulsion. Corresponding to these agents, there
may be recognized the following groups:
1. Water, hydrochores. These comprise all plants distributed exclusively
by water, whether the latter acts as ocean currents, tides, streams, or
surface run-off. In the case of streams and run-off, especially,
mobility plays little part, provided the disseminules are impervious or
little subject to injury by water. Motile plants, or those with motile
cells, which belong entirely to this group, may be distinguished as
autochores, which correspond closely to mastigospores.
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