Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
=282. Continuous and intermittent invasion.= With respect to the
frequency of migration, we may distinguish invasion as _continuous_, or
_intermittent_. Continuous invasion, which is indeed usually mutual,
occurs between contiguous formations of more or less similar character,
in which there is an annual movement from one into the other, and at the
same time a forward movement through each, resulting from the invaders
established the preceding year. By far the greater amount of invasion is
of this sort, as may readily be seen from the fact that migration varies
inversely as the distance, and ecesis may decrease even more rapidly
than the distance increases. The significant feature of continuous
invasion is that an outpost may be reinforced every year, thus making
probable the establishment of new outposts from this as a center, and
the ultimate extension of the species over a wide area. The
comparatively short distance and the regular alternation of migration
and ecesis render invasion of this sort very effective. An excellent
illustration of this is seen in transition areas and regions, which are
due directly to continuous and usually to mutual invasion. Intermittent
invasion results commonly from distant carriage, though it may occur
very rarely between dissimilar adjacent formations, when a temporary
swing in the physical factors makes ecesis possible for a time. It is
characterized by the fact that the succession of factors which have
brought about the invasion is more or less accidental and may never
recur. Intermittent invasion is relatively rare, and from the small
number of disseminules affected, it is of little importance in modifying
vegetation quantitatively. On the other hand, since a species may often
be carried far from its geographical area, it is frequently of great
significance in distribution.
=283. Complete and partial invasion.= When the movement of invaders into
a formation is so great that the original occupants are finally driven
out, the invasion may be termed _complete_. Such invasion is found
regularly in the case of many ruderal formations, and is typical of the
later stages of many successions. It is ordinarily the result of
continuous invasion. If the number of invaders is sufficiently small
that they may be adopted into the formation without radically changing
the latter, the invasion is _partial_. This is doubtless true of the
greater number of invasions, though these are regularly much less
striking and important than instances of complete invasion.
[Illustration: Fig. 59. Continuous invasion into a new area; mats of
_Arenaria sajanensis_. _Silene acaulis_ and _Sieversia turbinata_
invading an alpine gravel slide.]
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