Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
=293. Weathering.= Practically all primary successions start on soils
produced by weathering. This is also true of coral or volcanic islets
and of lava beds, for no terrestrial vegetation can secure a foothold
upon them until the surface of the rock has been to some extent
decomposed or disintegrated. Weathering, as is well known, consists of
two processes, disintegration and decomposition, which usually operate
successively, though they are sometimes concomitant. Disintegration
usually precedes, especially in rock masses, and unless it is soon
followed by decomposition, results in dysgeogenous soils. Decomposition
often goes hand in hand with disintegration, or it takes place so
rapidly and perfectly that it alone seems to be present. In either case,
the resulting soil is eugeogenous. The relation of decomposition to
disintegration determines the size and compactness of the soil
particles, and upon the latter depend the porosity, capillarity, and
hygroscopicity of the soil. These control in large degree the character
of the first vegetation to appear on the soil.
Another point of fundamental value in determining revegetation is the
disposition of the weathered rock. If it remains _in situ_, it will
evidently differ in respect to compactness, homogeneity,
nutrition-content, water-content, disseminules, etc., from weathered
material which has been transported. An essential difference also arises
from the fact that a rock may be weathered a long distance from the
place where the decomposed particles are finally deposited, and in the
midst of a vegetation very different from that found in the region of
deposit. The disposition of the weathered material affords in
consequence a satisfactory basis for the arrangement of primary
successions. The following classification is proposed, based upon the
soil groups established by Merrill.[39]
=294. Succession in residuary soils.= Residuary soils are always
sedentary, i. e., they are formed _in situ_. They show certain
differences dependent upon the rock from which they originate, which may
be mixed crystalline shale, sandstone, or limestone, but the
thoroughness of decomposition causes these differences to be
comparatively small. Residuary soils are typically eugeogenous; their
successions in consequence usually begin with mesophytes, and consist of
a few stages. The soluble salt-content is comparatively low, since all
soluble matters are readily leached out. Successions in these soils are
especially characteristic of shale, sandstone, and limestone ledges or
banks. Cumulose deposits, like residuary ones, are sedentary in
character, but as they are produced by the accumulation of organic
matter, they will be considered under reactions of vegetation upon
habitat.
[Illustration: Fig. 61. Talus arising from the disintegration of a
granitic cliff; the rocks are covered with crustose lichens.]
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