Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
=302. Succession by subsidence.= Subsidence is a factor of the most
profound importance in changing vegetation. It operates over vast areas
through immense periods of time. For these reasons, the changes are so
slow as to be almost imperceptible, and the resulting successions can be
studied only in the geological record. Extensive subsidence is confined
to-day to coastal plains, as in Greenland, the south Atlantic coast, and
the region of the Mississippi delta, where its effects are merged with
the paludation of tidal rivers, and the wave and tide erosion of the sea
shore. Such successions are unique, inasmuch as the denuding force
operates very slowly instead of quickly, and the first pioneers of the
new vegetation appear before the original formation has been destroyed.
In all cases, the succession is from mesophytic or halophytic formations
to paludose, and, finally, marine vegetation. In small areas of
subsidence, such as shore slips along lakes and streams, sink holes, and
sunken bogs, the succession is usually both short and simple, mesophytes
giving place to amphibious and ultimately to aquatic forms.
=303. Successions in landslips.= Landslips occur only in montane and
hilly regions, and here they are merely of local importance. In many
respects, they are not unlike talus; they show essential differences,
however, in that they are not sorted by gravity, and in that they
destroy vegetation almost instantly. The succession arises as a rule,
not upon the original soil, but upon that of the landslip, and, as
pointed out elsewhere, might well be regarded as primary.
=304. Succession in drained, or dried soils.= In geological times, the
subsidence of barriers must often have produced drainage and drying-out,
just as elevation frequently resulted in flooding and lake formation. At
the present time, the drying-out of lakes and ponds is the result of
artificial drainage, or of climatic changes. The former will be
considered under successions brought about by the agency of man.
Climatic changes when general operate so slowly that the stages of such
successions are perceptible only when recorded in strata. More locally,
climate swings back and forth through a period of years, with the result
that in dry years the swamps and ponds of wetter seasons are dried out,
and the vegetation destroyed or changed. If the process be gradual, the
succession passes from hydrophytic through amphibious to mesophytic,
and, in dry regions, xerophytic conditions. When the process of
drying-out occurs rapidly, as in a single summer, the original formation
is destroyed, and the new vegetation consists largely of ruderal plants.
A peculiar effect of climate occurs in regions with poor drainage, where
the result of intense evaporation is to produce alkaline basins and salt
lakes, in which the succession becomes more and more open, and is
finally represented by a few stabilized halophytes, or disappears
completely.
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