Research methods in ecologyClements, Frederic E. (Frederic Edward)
Science
Research methods in ecology
Clements, Frederic E. (Frederic Edward)
Plant ecology; Plant ecology -- Methodology
Single standards are exceedingly convenient in photometer readings, but
they are open to one objection. In the sunshine it is necessary to make
instant decision upon the accuracy of the match, or the exposure becomes
too deep. In the shade where the action is slower, this difficulty is
not felt. For this reason it is usually desirable to check the results
by a multiple standard, and in the case of selagraph records, where the
various exposures show a wide range of tint, light values are obtainable
only by direct comparison with the multiple standard. The exact matching
of exposure and standard requires great accuracy, but with a little
practice this may be done with slight chance of error by merely moving
the exposure along the various tints of the standard until the proper
shade is found. The requisite skill is soon acquired by running over a
strip of exposures several times until the comparisons always yield the
same results for each. The margin of error is practically negligible
when the same person makes all the comparisons, and in the case of two
or three working on the same reading the results diverge little or not
at all. The efficient difference for light is much more of a variable
than is the case with water-content. It has been determined so far only
for a few species, all of which seem to indicate that appreciable
modification in the form or structure of a leaf does not occur until the
reduction in intensity reaches .1 of the meridian sunlight at the June
solstice. The error of comparison is far less than this, and
consequently may be ignored, even in the most painstaking inquiry.
_Readings_
=85. Time.= The intensity of the light incident upon a habitat varies
periodically with the hour and the day, and changes in accord with the
changing conditions of the sky. The light variations on cloudy days can
only be determined by the photometer. While these can not be ignored,
proper comparisons can be instituted only between the readings taken on
normal days of sunshine. The sunlight varies with the altitude of the
sun, i. e., the angle which its rays make with the surface at a given
latitude. This angle reaches a daily maximum at meridian. The yearly
maximum falls on June 22, and the angle decreases in both directions
through the year to a minimum on December 22. At equal distances from
either solstice, the angle is the same, e. g., on March 21 and September
23. At Lincoln (41° N. latitude) the extremes at meridian are 73° and
26°; at Minnehaha (39°) they are 75° and 28°. The extremes for any
latitude may be found by subtracting its distance in degrees north of
the two tropics from 90. Thus, the 50th parallel is 26.5° north of the
tropic of Cancer, and the maximum altitude of the sun at a place upon it
is 63.5°. It is 73.5° north of the tropic of Capricorn, and the minimum
meridional altitude is 16.5°.
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