Researches on the Visual Organs of the TrilobitesLindström, Gustaf
Science
Researches on the Visual Organs of the Trilobites
Lindström, Gustaf
Trilobites
In fig. 2 a great change has set in. There is no distinction between
the lateral appendages of the third and fourth segments at left. These
two have been fused together, they have united, so as to make the left
triangular spine look larger than it was originally. The appendages
of the right side are still in the same state as before. But this
fusion of the lateral appendages also takes place in another direction,
as shown in another specimen (fig. 3), so that the second and third
appendages on both sides coalesce into one piece. Now it is easy to
imagine that at last a complete fusion has set in between all lateral
appendages and that instead of the original three on each side, there
is only one large piece, reaching beyond the shell as a broad spine, as
represented in the hypothetical figure 4.
In the figure 3 a progressive change is also seen in the transformation
of the first central segment. From occupying the whole foremost space
of the shell it has been lessened in size, more distantiated from the
anterior border of the shell and rounded off, more prominent and
definite from the lateral appendages, which have become narrow, though
of the same length as before. The two first central segments seem to
have been united into one.
Between the hypothetical figure 4 and fig. 5, there is evidently a
great lacuna, not yet filled up. In the interval of time the two
appendages, which we saw in fig. 4, have been much modified, the
posterior one having lost so much in bulk, and the anterior being
lengthened and stretching out beyond the border of the shell in a
narrow spine alongside the posterior one. The central segmented part
has now assumed a shape, which on comparing it with the following
stages of development makes its true nature evident and that it indeed
is what in the adult animal becomes the glabella. The meaning of the
previous stages then also is easily understood. The central segmental
piece in them is the glabella or we may, as BERNARD already has done,
call it for rhachis and the side appendages for pleura, as this little
larva represents the whole body of the future trilobite, and embodies
all its parts _in nuce_. Through the great changes which these pleura
undergo, it results, as we have seen, that two pair vanish, being
incorporated with the large fourth pair and that only two rest for a
while, the anterior one being the so much renowned frontal-lob or eye
lobe and the second one the so called »ornamental spine», which in fact
is a compound of the original second third and fourth pleura of the
corresponding segments.
It is to be borne in mind that this larva, which represents the future
trilobite in its earliest stage, is nothing but the head, or what
in the adult takes the place of the head, and especially its dorsal
surface and that it thus solely consists of the future head.
Public-domain text, read in full here on John Shaqi.
Reviews
Reviews
No reviews yet
Be the first to share your thoughts on this work.
Join the Discussion
Join the discussion
Sign in to leave a comment or review.
Sign InorCreate an account