Researches on the Visual Organs of the TrilobitesLindström, Gustaf
Science
Researches on the Visual Organs of the Trilobites
Lindström, Gustaf
Trilobites
appearance.
Towards both sides of the ocular surface, towards the superior and
inferior side, a change sets in as to the shape of the lenses, as best
seen in horizontal sections (figs. 8, 10) they are lengthened and
become more and more indistinct, and at last in the upper and lower
marginal zones pass over into a reticulate, spongious mass, which
seen in a longitudinal section presents almost the same aspect as in
the regular prismatic surface of the eye. Fig. 11 _b_, _a_ being the
visual field. It is, however, more densely reticulate, but a prismatic
arrangement is quite as much evident as in the ocular surface proper.
In the rule the passage from the prismatic surface to the reticulate
is gradual, the prisms becoming by and by irregular in their outline
and diminishing in size (fig. 12, _a_ the eye proper, _b_ the border
zone), but there are also instances where the distinction between these
two fields is sharp and without any gradual transition. In the inferior
reticulate zone there are generally some oblongue, funnelshaped pits.
I am uncertain whether they are to be regarded as regular parts of
the eye structure or rather as burrows of some parasite. They do not
continue deep down.
If we now compare the reticulate zone of the eye with the maculæ of the
hypostoma, for instance the vertical section fig. 16 with fig. 11 _b_,
we find the most complete identity in structure. In the same manner
the horizontal section of the border zone of the eye, fig. 8 _a_, fig.
10 _b_, and of the macula, fig. 17 are similar. The same chaotic,
spongious mass in both, with some tendency to form prisms more evident
in the longitudinal sections, where the same reticulate structure with
predominant white, longitudinal streaks is so palpable. That there is
a complete identity in structure between the two, the macula of the
hypostoma and the border zones of the eye, is as evident as anything
can be, but as to the functional identity or what this function may
have been it is difficult to decide anything with certainty, at least
it seems to me that the capacity of vision must in both have been far
more restricted than in the eye proper. They rather give the impression
as of rudimentary visual organs.
=Asaphus raniceps= DALM.
Pl. I f. 23-26.
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