Researches on the Visual Organs of the TrilobitesLindström, Gustaf
Science
Researches on the Visual Organs of the Trilobites
Lindström, Gustaf
Trilobites
When LILJEVALL saw this group of granules he was at once struck with
the perfect, outward similarity between them and the facets of the
compound eyes on the upper surface of the head of this same species,
and I fully concurred in the same view. It now became of importance to
ascertain whether the intimate structure also corresponded in both, and
microscopic sections were accordingly prepared. Although not so clear
as in other species of this genus a vertical section of a cephalic
eye (Pl. II fig. 21) shows a median row of black spots which compared
with other sections (Pl. II fig. 9) must be the centres of the badly
preserved lenses. This is confirmed by the horizontal section (Pl. II
fig. 20) where the well known image of the polyedric facets appeared.
In the same manner the granulated spot of the hypostoma shows in a
vertical section even more clearly (pl. II fig. 27) the elongate,
coherent lenses with a black centre and the polyedric form of the
lenses in a horizontal section (Pl. II fig. 26). The assumption of a
pair of small adventive eyes on the exterior side of the hypostoma
became thus based on the clear evidence of the perfect structural
agreement between them and the eyes of the head. But it was necessary
to strengthen this evidence through extended researches on the
hypostoma of as many trilobite genera as possible and also to study the
intimate structure of the cephalic eyes in as many genera as could be
accessible.
We shall now give first a general account of the structure of the eyes
in the trilobites as far as we have been able to study them, and then
proceed to describe the maculæ of the various genera, which we have
observed.
With regard to the presence or non existence of visual organs, we must
remark that a considerably greater number of genera than those which
unanimously have been regarded as blind, also are devoid of eyes though
they still by many authors are ranked amongst the oculate ones and we
then had better first to make a review of them and thus to eliminate
them from the number of the oculate species.
=Blind Trilobites.=
In the chaos of generic forms and in the great disagreement which
prevails as to the systematizing of the trilobites of the Cambrian
time, there is a thorough revision of them highly needed by a person
having access not only to the literature, but also to the original
specimens. It is almost impossible in the present state of things to
tell with any degree of certainty how many well established genera had
been living during that period. Hence the difficulty of fixing the
systematic names of many specimens the visual organs of which are to be
described.
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