abolish consciousness. Hence, it is a highly probable conclusion, that consciousness in man depends upon the integrity of the anterior division of the brain, while the middle and hinder divisions of the brain, and the rest of the nervous centres, have nothing to do with it. And it is further highly probable, that what is true for man is true for other vertebrated animals. We may assume, then, that in a living vertebrated animal, any segment of the cerebro-spinal axis (or spinal cord and brain) separated from that anterior division of the brain which is the organ of consciousness, is as completely incapable of giving rise to consciousness, as we know it to be incapable of carrying out volitions. Nevertheless, this separated segment of the spinal cord is not passive and inert. On the contrary, it is the seat of extremely remarkable powers. In our imaginary case of injury, the man would, as we have seen, be devoid of sensation in his legs, and would have not the least power of moving them. But, if the soles of his feet were tickled, the legs would be drawn up, just as vigorously as they would have been before the injury. We know exactly what happens when the soles of the feet are tickled; a molecular change takes place in the sensory nerves of the skin, and is propagated along them and through the posterior roots of the spinal nerves, which are constituted by them, to the grey matter of the spinal cord. Through that grey matter, the molecular motion is reflected into the anterior roots of the same nerves, constituted by the filaments which supply the muscles of the legs, and, travelling along these motor filaments, reaches the muscles, which at once contract, and cause the limbs to be drawn up. In order to move the legs in this way, a definite co-ordination of muscular contractions is necessary; the muscles must contract in a certain order and with duly proportioned force; and moreover, as the feet are drawn away from the source of irritation, it may be said that the action has a final cause, or is purposive. Thus it follows, that the grey matter of the segment of the man’s spinal cord, though it is devoid of consciousness, nevertheless responds to a simple stimulus by giving rise to a complex set of muscular contractions, co-ordinated towards a definite end, and serving an obvious purpose. If the spinal cord of a frog is cut across, so as to provide us with a segment separated from the brain, we shall have a subject parallel to the injured man, on which experiments can be made without remorse; as we have a right to conclude that a frog’s spinal cord is not likely to be conscious, when a man’s is not.