The tendency to produce rounded stems and leaves, little
distinguishable from one another, is equally noticeable in many seaside
plants which frequent the strip of thirsty sand beyond the reach of the
tides. That belt of dry beach that stretches between high-water mark
and the zone of vegetable mould, is to all intents and purpose a
miniature desert. True, it is watered by rain from time to time; but
the drops sink in so fast that in half an hour, as we know, the entire
strip is as dry as Sahara again. Now there are many shore weeds of this
intermediate sand-belt which mimic to a surprising degree the chief
external features of the cactuses. One such weed, the common
salicornia, which grows in sandy bottoms or hollows of the beach, has a
jointed stem, branched and succulent, after the true cactus pattern,
and entirely without leaves or their equivalents in any way. Still more
cactus-like in general effect is another familiar English seaside weed,
the kali or glasswort, so called because it was formerly burnt to
extract the soda. The glasswort has leaves, it is true, but they are
thick and fleshy, continuous with the stem, and each one terminating in
a sharp, needle-like spine, which effectually protects the weed against
all browsing aggressors.
Now, wherever you get very dry and sandy conditions of soil, you get
this same type of cactus-like vegetation--_plantes grasses_, as the
French well call them. The species which exhibit it are not necessary
related to one another in any way; often they belong to most widely
distinct families; it is an adaptive resemblance alone, due to
similarity of external circumstances only. The plants have to fight
against the same difficulties, and they adopt for the most part the
same tactics to fight them with. In other words, any plant of whatever
family, which wishes to thrive in desert conditions, must almost, as a
matter of course, become thick and succulent, so as to store up water,
and must be protected by a stout epidermis to prevent its evaporation
under the fierce heat of the sunlight. They do not necessarily lose
their leaves in the process; but the jointed stem usually answers the
purpose of leaves under such conditions far better than any thin and
exposed blade could do in the arid air of a baking desert. And
therefore, as a rule, desert plants are leafless.
Public-domain text, read in full here on John Shaqi.
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