Scientific American Supplement, No. 648, June 2, 1888.Various
Science
Scientific American Supplement, No. 648, June 2, 1888.
Various
Science -- Periodicals
The second generalization is, that since the cambium adds new elements to
the cortex on the inside of the latter, and since the cambium cylinder as
a whole is traveling radially outward--_i.e._, further from the pith--each
year, as follows from its mode of adding the new annual rings of wood on
to the exterior of the older ones, it is clear that the cortical jacket as
a whole must suffer distention from within, and tend to become too small
for the enlarging cylinder of rigid wood and growing cambium combined.
Indeed, it is not difficult to see that unless certain provisions are made
for keeping up the continuity of the cortical tissues, they must give way
under the pressure from within. As we shall see, such a catastrophe is in
part prevented by a very peculiar and efficient process.
Before we can understand this, however, we must take a glance at the
structural characters of the whole of this jacket (Fig. 20). While the
branch or stem is still young, it may be conveniently considered as
consisting of three chief parts.
(1) On the outside is a thin layer of flat, tabular cork cells (Fig. 20,
C_o_), which increase in number by the activity of certain layers of cells
along a plane parallel to the surface of the stem or branch. These cells
(C.C_a_) behave very much like the proper cambium, only the cells divided
off from them do not undergo the profound changes suffered by those which
are to become elements of the wood and inner cortex. The cells formed on
the outside of the line C.C_a_ in fact simply become cork cells; while
those formed on the inside of the line C.C_a_ become living cells (C_l_)
very like those I am now going to describe.
(2) Inside this cork-forming layer is a mass of soft, thin-walled "juicy"
cells, _pa_, which are all living, and most of which contain granules of
chlorophyl, and thus give the green color to the young cortex--a color
which becomes toned down to various shades of olive, gray, brown, etc., as
the layers of cork increase with the age of the part. It is because the
corky layers are becoming thicker that the twig passes from green to gray
or brown as it grows older. Now, these green living cells of the cortex
are very important for our purpose, because, since they contain much food
material and soft juicy contents of just the kind to nourish a parasitic
fungus, we shall find that, whenever they are exposed by injury, etc.,
they constitute an important place of weakness--nay, more, various fungi
are adapted in most peculiar ways to get at them. Since these cells are
for the most part living, and capable of dividing, also, we have to
consider the part they play in increasing the extent of the cortex.
Public-domain text, read in full here on John Shaqi.
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