Scientific American Supplement, No. 648, June 2, 1888. — John Shaqi
Scientific American Supplement, No. 648, June 2, 1888.Various
Science
Scientific American Supplement, No. 648, June 2, 1888.
Various
Science -- Periodicals
The above brief account may suffice to give a general idea of what the
cortical jacket covering our timber is, and how it comes about that in the
normal case the thickening of the cylinder is rendered possible without
exposing the cambium and other delicate tissues: it may also serve to show
why bark is so various in composition and other characters. But it is also
clear that this jacket of coherent bark, bound together by the elastic
sheets of cork, must exert considerable pressure as it reacts on the
softer, living, succulent parts of the cortex, trapped as they are between
the rigid wood cylinder and the bark; and it is easy to convince
ourselves that such is the case. By simply cutting a longitudinal slit
through the cortex, down to near the cambium, but taking care not to
injure the latter, the following results may be obtained. First, the bark
gapes, the raw edges of the wound separating and exposing the tissues
below; next in course of time the raw edges are seen to be healed over
with cork--produced by the conversion of the outer cells into cork cells.
As time passes, provided no external interference occurs, the now rounded
and somewhat swollen cork-covered edges of the wound will be found closing
up again; and sooner or later, depending chiefly on the extent of the
wound and the vigor of the tree, the growing lips of the wound will come
together and unite completely.
But examination will show that although such a slit wound is so easily
healed over, it has had an effect on the wood. Supposing it has required
three years to heal over, it will be found that the new annual rings of
wood are a little thicker just below the slit; this is simply because the
slit had released the pressure on the cambium. The converse has also been
proved to be true--_i.e._, by increasing the pressure on the cambium by
means of iron bands, the annual rings below the bands are thinner and
denser than elsewhere.
But we have also seen that the cambium is not the only living tissue below
the bark: the cortical parenchyma (_pa_) and the cells (_c_) of the inner
cortex (technically the phloem) are all living and capable of growth and
division, as was described above. The release from pressure affects them
also; in fact, the "callus," or cushion of tissue which starts from the
lips of the wound and closes it over, simply consists of the rapidly
growing and dividing cells of this cortex, _i.e._, the release from
pressure enables them to more than catch up the enlarging layer of cortex
around the wound.
An elegant and simple instance of this accelerated growth of the cortex
and cambium when released from the pressure of other tissues is exhibited
in the healing over of the cut ends of a branch, a subject to be dealt
with later on; and the whole practice of propagation by slips or cuttings,
the renewal of the "bark" of cinchonas, and other economic processes,
depend on these matters.
Public-domain text, read in full here on John Shaqi.
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