The eggs are not always laid in the condition of the simple planula
described above. They may, on the contrary, be dropped from the parent
in different stages of development, sometimes even after the tentacles
have begun to form, as in Figs. 8, 9. Neither is it by means of eggs
alone that these animals reproduce themselves; they may also multiply
by a process of self-division. The disk of an Actinia may contract
along its centre till the circular outline is changed to that of a
figure 8, this constriction deepening gradually till the two halves of
the 8 separate, and we have an Actinia with two mouths, each
surrounded by an independent set of tentacles. Presently this
separation descends vertically till the body is finally divided from
summit to base, and we have two Actiniæ where there was originally but
one. Another and a far more common mode of reproduction among these
animals is that of budding like corals. A slight swelling arises on
the side of the body or at its base; it enlarges gradually, a
digestive cavity is formed within it, tentacles arise around its
summit, and it finally drops off from the parent and leads an
independent existence. As a number of these buds are frequently formed
at once, such an Actinia, surrounded by its little family, still
attached to the parent, may appear for a time like a compound stock,
though their normal mode of existence is individual and distinct.
The Actinia is exceedingly sensitive, contracting the body and drawing
in the tentacles almost instantaneously at the slightest touch. These
sudden movements are produced by two powerful sets of muscles, running
at right angles with each other through the thickness of the body
wall; the one straight and vertical, extending from the base of the
wall to its summit; the other circular and horizontal, stretching
concentrically around it. By the contraction of the former, the body
is of course shortened; by the contraction of the latter, the body is,
on the contrary, lengthened in proportion to the compression of its
circumference. Both sets can easily be traced by the vertical and
horizontal lines crossing each other on the external wall of the body,
as in Fig. 5. Each tentacle is in like manner furnished with a double
set of muscles, having an action similar to that described above. In
consequence of these violent muscular contractions, the water imbibed
by the animal, and by which all its parts are distended to the utmost,
is forced, not only out of the mouth, but also through small openings
in the body wall scarcely perceptible under ordinary circumstances,
but at such times emitting little fountains in every direction.
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