[Illustration: Fig. 154. Magnified view of the group of young
Comatulæ of Fig. 153.]
We are indebted to Thompson for the explanation of the true relations
of the young Comatula to the present Pentacrinus and the fossil
Crinoids. Supposing these young to be full-grown animals, he at first
described them as living representatives of the genus Pentacrinus; it
was only after he had watched their development, and ascertained by
actual observation that they dropped from their stem, to lead an
independent life as free Comatulæ, that he fully understood their true
connection with the past history of their kind, as well as with their
contemporaries. In Fig. 153, a faint star-like dot (_y_) may be seen
attached to the side of the disk by a slight line. In Fig. 154, we
have that minute dot as it appears under the microscope, magnified
many diameters; when it is seen to be a cirrus of a Comatula, with
three small Pentacrinus-like animals growing upon it, in different
stages of development. In the upper one, the branching arms and the
disk, with its many plates, are already formed; and though in the
figure the rays are folded together, they are free, and can be opened
at will. In the larger of the two lower buds, the plates of the disk
are less perfect, and the arms are straight and simple, without any
ramifications, though they are free and movable, whereas, in the
smaller one, they are folded within the closed bud.
* * * * *
EMBRYOLOGY OF ECHINODERMS.
All Radiates have a special mode of development, as distinct for each
class as is their adult condition, and in none are the stages of
growth more characteristic than in the Echinoderms. In the Polyps, the
division of the body into chambers, so marked a feature of their
ultimate structure, takes place early; in the Acalephs, the tubes
which traverse the body are hollowed out of its mass in the first
stages of the embryonic growth, and we shall see that in the
Echinoderms also, the distinctive feature of their structure, viz. the
enclosing of the organs by separate walls, early manifests itself.
This peculiarity gives to the internal structure of these animals so
individual a character, that some naturalists, overlooking the law of
radiation, as prevalent in them as in any members of this division,
have been inclined to separate them, as a primary division of the
animal kingdom, from the Polyps and Acalephs, in both of which the
body-wall furnishes the walls of the different internal cavities,
either by folding inwardly in such a manner as to enclose them, as in
the Polyps, or by the cavities themselves being hollowed out of the
general mass, as in the Acalephs.
_Star-fish_. (_Astracanthion_.)
[Illustration: Fig. 155. Egg of Star-fish.]
[Illustration: Fig. 156. Egg of Star-fish in which the yolk has
been divided into two segments.]
[Illustration: Fig. 157. Egg in which there are eight segments of
the yolk.]
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