The egg of the Star-fish, when first formed, is a transparent,
spherical body, enclosing the germinative vesicle and dot. (See Fig.
155.) As soon as these disappear, the segmentation of the yolk begins;
it divides first into two portions (see Fig. 156), then into four,
then into eight, and so on; but when there are no more than eight
bodies of segmentation (see Fig. 157), they already show a disposition
to arrange themselves in a hollow sphere, enclosing a space within,
and by the time the segmentation is completed, they form a continuous
spherical shell. At this time the egg, or, as we will henceforth call
it, the embryo, escapes and swims freely about. (See Fig. 158.) The
wall next begins to thin out on one side, while on the opposite side,
which by comparison becomes somewhat bulging, a depression is formed
(_m_ _a_, Fig. 159), gradually elongating into a loop hanging down
within the little animal, and forming a digestive cavity. (_d_, Fig.
160.) At this stage it much resembles a young Actinia. The loop
spreads somewhat at its upper extremity, and at its lower end is an
opening, which at this period of the animal's life serves a double
purpose, that of mouth and anus also, for at this opening it both
takes in and rejects its food. We shall see that before long a true
mouth is formed, after which this first aperture takes its place
opposite the mouth, retaining only the function of the anus. Presently
from the upper bulging extremity of the digestive cavity, two lappets,
or little pouches, project (_w_ _w'_ Fig. 161); they shortly become
completely separated from it, and form two distinct hollow cavities
(_w_ _w'_, Fig. 162). Here begins the true history of the young
Star-fish, for these two cavities will develop into two water-tubes,
on one of which the back of the Star-fish, that is, its upper surface,
covered with spines, will be developed, while on the other, the lower
surface, with the suckers and tentacles, will arise. At a very early
stage one of these water-tubes (_w'_, Fig. 163) connects with a
smaller tube opening outwards, which is hereafter to be the madreporic
body (_b_, Fig. 163). Almost until the end of its growth, these two
surfaces, as we shall see, remain separate, and form an open angle
with one another; it is only toward the end of the development that
they unite, enclosing between them the internal organs, which have
been built up in the mean while.
[Illustration: Fig. 158. Larva just hatched from egg; a thickened
pole.]
[Illustration: Fig. 159. Larva somewhat older than Fig. 158; _m_
_a_ depression at thickened pole.]
[Illustration: Fig. 160. Larva where the depression has become a
digestive cavity _d_, opening at _a_.]
[Illustration: Fig. 161. Earlets, _w_ _w'_ (water-tubes),
developed at the extremity of the digestive cavity _d_; _m_
mouth.]
[Illustration: Fig. 162. More advanced larva; _a_ _d_ _c_
digestive system, _v_ vibratile chord, _m_ mouth.]
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