It was, moreover, during his official connection with the Middlesex Hospital that Mr. Flower married Georgina Rosetta, the youngest daughter of the late Admiral W. H. Smyth, C.S.I., etc., a well-known astronomer, who was for some time Hydrographer to the Admiralty and likewise Foreign Secretary to the Royal Society, the wedding taking place in 1858 at the church of Stone, in Buckinghamshire, near the bride’s home. This happy union had in many ways an important influence upon the future career of the young surgeon, for, in addition to her father, several of the relatives of Mrs. (now Lady) Flower were more or less intimately connected with scientific work and scientific people; among them being Sir Warrington Smyth (sometime Inspector-General of Mines), Professor Piazzi Smyth, General Sir Henry Smyth, and Sir George Baden-Powell. It was to Lady Flower that Sir William dedicated his last work, the volume entitled _Essays on Museums_. A tour through Belgium and up the Rhine followed the marriage. Although it scarcely comes within the purview of this biography to allude to the issue of this marriage, it may be mentioned that of the three sons born to Sir William Flower, the second alone, Stanley Smyth, inherited his father’s zoological tastes. Captain S. S. Flower (who takes his first name from Dean Stanley, of Westminster, an intimate friend of the family), after serving for some time in the 5th Fusileers, obtained the appointment of Director of the Royal Museum at Bangkok, Siam, after which he was made Director of the Khedival Zoological Gardens at Giza, near Cairo, to which post (which he still holds) was subsequently added that of Superintendent of Game Protection in the Sudan. Captain Flower has not only raised the menagerie at Giza to a high state of perfection, but has contributed several papers to the _Proceedings_ of the Zoological Society of London on the zoology of Siam and the Malay countries. To revert to the proper subject of this memoir, during his tenure of the aforesaid official posts at the Middlesex Hospital it was apparent to his intimate scientific friends—among whom were included the late Professor T. H. Huxley and the late Mr. George Busk—that the inclinations of Flower were all on the side of comparative anatomy rather than towards practical surgery or medicine. Accordingly, when the appointment of Conservator to the Museum of the Royal College of Surgeons became vacant in 1861 by the death of Mr. Quekett, Flower was strongly recommended by Huxley (then Hunterian Professor), Busk, and other friends as a suitable successor, and was in due course elected by the Council. When, nine years later (1870), Huxley himself felt compelled by the pressure of other engagements and work to resign the Hunterian Professorship, the Conservator of the Museum was appointed to the vacant chair, thus once more bringing together two posts which had been sundered since Owen’s resignation.