Speciation in the Brazilian Spiny Rats — John Shaqi
Speciation in the Brazilian Spiny RatsMoojen, João
Science
Speciation in the Brazilian Spiny Rats
Moojen, João
Proechimys
_Diagnosis._--Upper parts Mars Orange on back, grading to
Ochraceous-Tawny on sides; zygomatic breadth narrow; nasals short;
incisive foramen narrow and short; vomerine sheath complete and
thick; upper molars usually with four counterfolds.
_Pelage._--_Aristiforms on middorsal region_: Grayish basally,
gradually blackening toward tip; total length, 16 to 19 mm;
maximum width, 0.5 mm. _Setiforms on middorsal region_: Grayish on
basal third, gradually blackening toward tip but interrupted by a
Mars Orange, subapical zone 1.5 mm long; total length, 16 to 19
mm; maximum width, 0.06 mm. _Setiforms on outer thighs_: Whitish
basally, gradually blackening toward tip but interrupted by Orange
Rufous or Ochraceous-Tawny, subapical zone; total length, 14 to 16
mm; maximum width, 0.05 mm.
[Illustration: FIGS. 25, 27. _Proechimys goeldii steerei_, sex ?,
"Hyutanaham," USNM no. 105537. × 1.]
[Illustration: FIGS. 26, 28. _Proechimys goeldii goeldii_, female, AMNH
no. 37488. × 1.]
[Illustration: FIGS. 29, 30. _Proechimys goeldii steerei_, sex ?, USNM
no. 105537, "Hyutanaham." × 1.]
[Illustration: FIGS. 31, 32. _Proechimys goeldii goeldii_, female, AMNH
no. 37488, Fazenda Paraiso. × 1.]
_Skull._--Large and strong; rostrum rather pointed posteriorly;
supraorbital ridges not much expanded and extending across
anterior half of parietals; infraorbital foramen without groove
for transmission of nerve, or groove obsolete; zygomatic arches
slender; postorbital process of zygoma involving mostly squamosal;
incisive foramen short and narrow (4.5 × 2.5 mm) with margins
almost parallel or tapering gradually caudad and extending toward
palate as ridges; posterior margin of incisive foramen
approximately 2.5 mm anterior to premolars; vomerine sheath
complete, with both elements well-developed; mesopterygoid fossa
never extending anterior to middle of M3; bullae large, well
inflated and with shallow grooves.
_Teeth._--Upper molariform teeth: P4 with three counterfolds;
upper molars with four counterfolds each or, less commonly, three.
Lower premolars with four counterfolds; lower molars with three
each.
_Comparisons._--From _P. g. goeldii_, _steerei_ differs in: Back
and sides with more reddish; narrower interorbitally and across
zygomata; palatilar length less and nasals shorter; maxillary part
of vomerine sheath thicker; usually four instead of three
counterfolds in M3.
_Remarks._--This subspecies is clearly related to _P. goeldii_. One
skull from Porto Velho, Rio Madeira, _Guaporé_, Brazil (CNHM no. 21558)
may belong to an unnamed subspecies but is provisionally included here.
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