Speciation in the Brazilian Spiny RatsMoojen, João
Science
Speciation in the Brazilian Spiny Rats
Moojen, João
Proechimys
_Teeth._--Molariform teeth large, P4-M3 averaging more than 9 mm
in length. Upper molariform teeth: P4 and M3 with three
counterfolds; M1 and M2 with four counterfolds each. In lower
teeth, p4 with four counterfolds and each molar with three
counterfolds.
_Comparisons._--Differences from _P. g. steerei_ are given in the
account of that subspecies.
_Remarks._--Specimens from the type locality were not available.
Specimens from Fazenda Paraiso, Faro, were relied upon as representative
of the subspecies. These agree with the type according to Thomas
(1912:89). However, the skin of the type was changed in color by
preservative (Thomas, 1905:587) and the best skin he saw was from Faro
(1912:89).
Thomas (1920:277) applied the name _goeldii_ also to specimens from
Manacaparú, a place a short distance above Manaus on the Solimões
(Amazon) River and from Acajutuba, near Manaus, on the Negro River. In
referring to these specimens (2 from Manacaparú and 2 from Acajutuba)
Thomas (_loc. cit._) said "Five molar laminae are frequently, if not
invariably, present among these specimens." He did not, however, mention
whether or not the number of laminae was constant in both M2 and M3. One
specimen from Acajutuba, in the collection of Museu Nacional (MN no.
1973 [M]), actually has five laminae in M3, but the specimens in the
American Museum from Faro agree absolutely with Thomas' original
description of _goeldii_.
Osgood (1944:199) doubted that _goeldii_ was a valid species. Evidence
that Osgood's doubt was unjustified is furnished by the fact that Thomas
(1912:89) pointed out that his specimen from Faro agrees with the type.
Likewise, my two specimens from Faro agree with the type insofar as it
has been described. Thomas (1912:89) mentioned two additional skulls
from the type locality which, he stated, agree with the type which was
received from the Museu Goeldi, Pará.
_Specimens examined._--Total number, 4, from Brazil as follows:
Pará, _Faro_, Faro, Fazenda Paraiso, 2 (AMNH); Amazonas, _Manaus_,
Manaus, 1 skull (AMNH); Amazonas, _Manaus_, Acajutuba, 1 (MN).
_Additional records._--Total number, 7 (British Museum), from
Brazil, as follows: Pará, _Santarem_, Santarem (Thomas, 1912:89;
1920:277), 3; Amazonas, _Manaus_, Acajutuba (Thomas, 1920:277), 2;
_Manacaparú_, Manacaparú (Thomas, 1920:277), 2.
=Proechimys semispinosus= (Tomes)
[Illustration: FIGS. 33, 36. _Proechimys semispinosus liminalis_,
female, MN no. 6253, Rio Quichito. Type. × 1.]
[Illustration: FIGS. 34, 37. _Proechimys semispinosus amphichoricus_,
male, AMNH no. 77020, Mount Duida. Type. × 1.]
[Illustration: FIGS. 35, 38. _Proechimys semispinosus kermiti_, female,
AMNH no. 37124, Lower Rio Solimões. Type. × 1.2 (from photograph).]
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