Speciation in the Brazilian Spiny RatsMoojen, João
Science
Speciation in the Brazilian Spiny Rats
Moojen, João
Proechimys
_Pelage._--_Aristiforms on middorsal region_: Grayish basally,
gradually blackening toward tip; total length, 18 to 20 mm;
maximum width, 0.8 to 1.0 mm. _Setiforms on middorsal region_:
Gray basally, gradually blackening toward tip but interrupted by a
light (16 i), Buckthorn Brown, subapical zone 2 mm long; total
length, 18 to 22 mm; maximum width, 0.03 mm. Most of them,
however, whitish basally, gradually blackening toward tip without
any distinctively-colored, subapical zone; total length, 24 to 26
mm; maximum width, 0.5 mm. _Setiforms on outer thighs_: Whitish
basally, gradually blackening toward tip but interrupted by an
Ochraceous-Buff, subapical zone 3.5 mm long; black tip short; total
length, 17 to 19 mm; maximum width, 0.05 mm.
_Skull._--Large and slender; rostrum elongate; nasals bluntly pointed
posteriorly; supraorbital ridges thick (but not expanded) and extending
across parietals but almost obsolete in middle part of parietals;
infraorbital foramen with weakly-developed groove for transmission of
nerve; postorbital process of zygoma involving mostly squamosal;
incisive foramen 5.5 x 2.8 mm wide in anterior third, with margins
constricted posteriorly and extending as ridges approximately 2 mm
beyond posterior margin of incisive foramen; posterior margin of
incisive foramen approximately 2.5 mm anterior to premolars; vomerine
sheath complete with maxillary part weak and premaxillary part extending
posteriorly beyond middle of incisive foramen; mesopterygoid fossa
extending forward as far as middle of M3; bullae well inflated and
elongated.
_Teeth._--P4 with four counterfolds in one of five specimens and with
three in remainder; M1 with four counterfolds in three of five specimens
and with three in remainder; M2 with three counterfolds in one specimen
and with four in all four remaining specimens; M3 always with four
counterfolds. Lower premolars with four counterfolds and lower molars
with only three.
_Comparisons._--The subspecies is easily distinguishable from _P. s.
angularis_ by: larger number of black setiforms on back, forming an
almost black longitudinal band; more elongate skull; larger and longer
bulla; longer incisive foramen which is more constricted posteriorly.
_Specimens examined._--Total number, 6 (AMNH), as follows:
Venezuela, territ. _Amazonas_, Esmeralda, Mt. Duida, altitude 325
m., 4; Venezuela, territ. _Amazonas_, Rio Cassiquiare, Quemapuré,
1; Brazil, Amazonas, _São Gabriel_, Rio Uaupés or Caiari, Tatú, 1.
=Proechimys semispinosus kermiti= Allen
_Proechimys kermiti_ Allen, 30 December 1915, Bull. Amer. Mus. Nat.
Hist., 34(22):629 (orig. descr.); Allen, 1916, Bull. Amer. Mus.
Nat. Hist., 35(30):569; Tate, 1935, Bull. Amer. Mus. Nat. Hist.,
68(5):400; Ellerman, 1940, The families and genera of living
rodents, Brit. Mus. (Nat. Hist.), 1:119.
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