Speciation of the Wandering ShrewFindley, James S. (James Smith)
Science
Speciation of the Wandering Shrew
Findley, James S. (James Smith)
Shrews
_Comparisons._--For comparison with _S. v. monticola_ see account of
that subspecies. Differs from _S. v. halicoetes_ in relatively
narrower and more attenuate rostrum and in less brownish underparts in
winter pelage; smaller and more brownish (less grayish) than _Sorex
vagrans_ from the southern Sierra Nevada.
[Illustration: FIG. 18. Probable geographic ranges of _Sorex
vagrans vagrans_, its derivative subspecies, and _S. v.
mixtus_.
1. _S. v. vancouverensis_
2. _S. v. vagrans_
3. _S. v. halicoetes_
4. _S. v. paludivagus_
5. _S. v. obscuroides_
6. _S. v. mixtus_
]
_Remarks._--Restriction of the range of _S. v. monticola_ to Arizona
and New Mexico leaves shrews that were formerly assigned to this
subspecies from Utah, Idaho, Washington and southern British Columbia
unassigned. Several names are available for consideration. The name
_Sorex vagrans dobsoni_ Merriam, 1891, type locality Alturas Lake,
Blaine Co., Idaho, was once applied to small shrews from Idaho,
Montana, Wyoming, and Utah, but was considered by Jackson to be
synonymous with _S. v. monticola_. The name _Sorex vagrans amoenus_
Merriam, 1895, type locality near Mammoth, Mono Co., California, has
been applied to wandering shrews from western Nevada, northeastern
California and southern Oregon. _Sorex vagrans nevadensis_ Merriam,
1895, type locality Reese River on Nye-Lander Co. line, Nevada was
considered by Hall (1946:119) to be synonymous with _S. v. amoenus_.
Specimens of _Sorex vagrans_ west of the Cascade Mountains have long
been referred to the nominate subspecies which has its type locality
at Willapa Bay, Pacific Co., Washington. Over so wide an area it is
only to be expected that some geographic variation is to be found.
Thus specimens from central Nevada average slightly paler in summer
pelage than those from the Pacific Coast or from the foothills of the
Rocky Mountains. In addition there are slight average differences in
size from place to place. Topotypes of _S. v. vagrans_, however, show
a fair degree of variability and some are nearly as pale as the paler
Great Basin stocks. Furthermore topotypical individuals of _vagrans_
can be lost in series of _S. v. amoenus_, although _amoenus_ is
shorter-tailed on the average. Specimens from the western foothills
of the Rocky Mountains show an amazing series of relationships with
the montane _S. v. obscurus_. In Utah, as previously pointed out,
complete intergradation occurs. At 1 mi. N Heath, Washington Co.,
Idaho, the lowland and the highland forms approach each other within a
short distance and still maintain a degree of distinctness, especially
in size. In northwestern Montana intergradation is extensive
(Clothier, 1950). In northeastern Washington distinctly separable
populations occur within a few miles of one another. In southern
British Columbia some populations are clearly intergrades while at 6
mi. S Yahk intergradation seemingly has not taken place. Where some
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