About the year 1840, this corn was introduced into Baden in Germany, and
cultivated by Metzger. In the first year it came true to type, and [206]
attained a height of 12 feet, but the season did not allow its seeds to
ripen normally. Only a few kernels were developed before the winter.
From this seed plants of a wholly different type came the next year, of
smaller stature, and with more brownish and rounded kernels. They also
flowered earlier and ripened a large number of seeds. The depression on
the outer side of the seed had almost disappeared, and the original
white had become darker. Some of the seeds had even become yellow and in
their rounded form they approached the common European maize. Obviously
they were hybrids, assuming the character of their pollen-parent, which
evidently was the ordinary corn, cultivated all around. The observation
of the next year showed this clearly, for in the third generation nearly
all resemblance to the original and very distinct American species was
lost. If we assume that only those seeds ripened which reverted to the
early-ripening European type, and that those that remained true to the
very late American variety could not reach maturity, the case seems to
be wholly comprehensible, without supposing any other factors to have
been at work than those of vicinism, which though unknown at the period
of Metzger's and Darwin's writings, seems now to be fully understood. No
innate tendency to run out and no changing influence of the climate are
required for an adequate explanation of the facts.
In the observation quoted, what astonishes us most, is the great
rapidity of the change, and the short time necessary for the offspring
of the accidental crosses to completely supplant the introduced type. In
the lecture on the selection of elementary species, closely analogous
cases were described. One of them was the wild oat or _Avena fatua_
which rapidly supplants the cultivated oats in bad years in parts of the
fields. Other instances were the experiments of Risler with the
"Galland" wheat and the observation of Rimpau on "Rivett's bearded"
wheat.
Before leaving the question of vicinism and its bearing on the general
belief of the instability of varieties, which when tested with due care,
prove to be quite stable, it may be as well to consider the phenomena
from another point of view. Our present knowledge of the effects of
crosses between varieties enables us to formulate some general rules,
which may be used to calculate, and in some way to predict, the nature
of the impurities which necessarily attend the cultivation of allied
species in close vicinity. And this mode of cultivation being in almost
universal use in the larger nurseries, [208] we may, by this discussion,
arrive at a more scientific estimation of the phenomena of vicinism,
hitherto described.
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