The simplest case that may be given, is when an ordinary retrograde
variety is cultivated with the species to which it belongs. For
instance, if dwarfs are cultivated next to the taller type, or a white
variety next to the red or blue-flowering species, or thornless forms in
neighboring beds with the armed species. Bees and Bumble-bees,
butterflies and moths are seen flying from flower to flower, collecting
the honey and carrying pollen. I frequently saw them cross the limits of
the neighboring beds. Loaded with the pollen of the variety they visit
the flowers of the different species and impregnate the stigma with it.
And returning to the variety they bring about similar crosses in the
flowers of the latter. Hybrid seeds will develop in both cases and
become mixed with the crop. We now have to ask the question, what sort
of plants will arise from these hybrid seeds. As a general rule we may
state, first, that the hybrids of either form of cross are practically
the same, secondly that they are not intermediate, but that the
character of one parent prevails to the almost absolute exclusion of the
other and in the third place that the older character dominates the
younger.
[209] The hybrid offspring will therefore, in the main, have the
character of the species and be indistinguishable from it, or show only
such differences as escape ordinary observation. When occurring in the
seeds of the variety they betray themselves as soon as the differential
characters are displayed. Between the thousands of flowering plants of a
white variety the hybrids will instantly catch the eye by their red or
blue corollas. Quite the contrary effect results from the admixture of
hybrids with the seeds of the species itself. Here no difference will
show itself, even in the fullest bloom. The effect of the spontaneous
crosses will pass unobserved. The strain, if pure in the first year,
will seem to be still in the same condition. Or in other terms, the
unavoidable spontaneous crosses will disturb the purity of the variety
in the second year, while they do not seem to interfere at all with the
uniformity of the species. The direct effect of the visits of the
insects is evident in the first case, but passes unobserved in the
latter.
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