From this it would seem, that spontaneous crosses are hurtful to
varieties, but are innocuous to true species. Certainly this would be
so, were there no selection. But it is easily seen, that through this
operation the effect becomes quite the opposite. For when the fields
[210] are inspected at the time of the fullest display of the varietal
characters, the obvious hybrids will be eliminated, but the hidden ones
will of necessity be spared, as they are concealed among the species by
the similarity of their type. Hence, the harvest of the variety may be
rendered pure or nearly so, while the harvest of the species will retain
the seeds of the hybrids. Moreover it will contain seeds originated by
the spontaneous but numerous crosses of the true plants with the
sparsely intermingled hybrids.
This brings us to the question, as to what will be the visible
consequences of the occurrence of such invisible hybrids in the
following generation. In opposition to the direct effects just
described, we may call them indirect. To judge of their influence, we
must know how hybrid seeds of the first generation behave.
In one of our lectures we will deal with the laws that show the
numerical relations known as the laws of Mendel. But for our present
purpose, these numerical relations are only of subordinate importance.
What interests us here is the fact that hybrids of varieties do not
remain constant in the second generation but usually split as it is
said, remaining hybrid only in part of their offspring, the other
portion returning to the parental types. This however, will show itself
only in those individuals [211] which reassume the character of the
varietal parent, all the others apparently remaining true to the type of
the species. Now it is easy to foresee what must happen in the second
generation if the first generation after the cross is supposed to be
kept free from new vicinistic influences, or from crosses with
neighboring varieties.
We may limit ourselves in the first place to the seeds of the unobserved
hybrids. For the greater part they will repeat the character of their
parents and still remain concealed. But a small number will display the
varietal marks, as for example showing white flowers in a field of blue
ones. Hence, the indirect consequence of the spontaneous crosses will be
the same in the species, as was the direct effect in the variety, only
that it appears a year later. It will then be eliminated in the process
of selection.
Obviously, this elimination conduces only to a partial purification. The
conspicuous plants will be destroyed, but a greater number of hybrids
will remain, still concealed by their resemblance to the general type
and will be spared to repeat the same process next year. So while the
variety may be freed every year from the impurities brought into it in
the preceeding summer, the admixtures of the species [212] will continue
during a number of years, and it may not be possible to get rid of them
at all.
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