Leaving this first generation, the original hybrids, we now come to a
discussion of their offspring. Hybrids should be fertilized either by
their own pollen, or by that of other individuals born from the same
cross. Only in this case can the offspring be considered as a means of
arriving at a decision as to the internal nature of the hybrids
themselves. Breeders generally prefer to fertilize hybrids with the
pollen of their parents. But this operation is to be considered as a new
cross, and consequently is wholly excluded from our present discussion.
Hence it follows that a clear insight into the heredity of hybrids may
be expected only from scientific experiments. Furthermore some of the
diversity observed as a result of ordinary crosses, may be due to the
instability of the parents themselves or at least of one of them, since
breeders ordinarily choose for their crosses some already very variable
strain. Combining such a strain with the desirable qualities of some
newly imported species, a new strain may [287] result, having the new
attribute in addition to all the variability of the old types. In
scientific experiments made for the purpose of investigating the general
laws of hybridity, such complex cases are therefore to be wholly
excluded. The hereditary purity of the parents must be considered as one
of the first conditions of success.
Moreover the progeny must be numerous, since neither constancy, nor the
exact proportions in the case of instability, can be determined with a
small lot of plants.
Finally, and in order to come to a definite choice of research material,
we should keep in mind that the chief object is to ascertain the
relation of the offspring to their parents. Now in nearly all cases the
seeds are separated from the fruits and from one another, before it
becomes possible to judge of their qualities. One may open a fruit and
count the seeds, but ordinarily nothing is noted as to their characters.
In this respect no other plant equals the corn or maize, as the kernels
remain together on the spike, and as it has more than one variety
characterized by the color, or constitution, or other qualities of the
grains. A corn-grain, however, is not a seed, but a fruit containing a
seed. Hence the outer parts pertain to the parent plant and only the
innermost ones to the [288] seedling and therefore to the following
generation. Fruit-characters thus do not offer the qualities we need,
only the qualities resulting from fertilizations are characteristic of
the new generation. Such attributes are afforded in some cases by the
color, in others by the chemical constitution.
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