Very few generations of such opposite selection are required to reduce
the race to an utterly poor one. In three years I was able to nearly
obliterate the type of my variety. I chose the seedlings with an
undivided primary leaf, cultivated them and counted their offspring
separately after the sowing. I found some parents with only 2-3% of
seedlings with divided primary leaves. And by a repeated selection in
this retrograde direction I succeeded in getting a great number of
plants, which during the whole summer made only very few leaves with
more than three blades. But an absolute reversion could no more be
reached in this direction than in the normal one. Any sowing without
selection would be [352] liable to reduce the strain to an average
condition.
The production of varietal and of atavistic leaves is dependent to a
high degree on external conditions. It agrees with the general rule,
that favorable circumstances strengthen the varietal peculiarities,
while unfavorable conditions increase the number of the parts with the
atavistic attribute. These influences may be seen to have their effect
on the single individuals, as well as on the generations growing from
their seed. I cannot cite here all the experimental material, but a
single illustrative example may be given. I divided a strong individual
into two parts, planted one in rich soil and the other in poor sand, and
had both pollinated by bees with the pollen of some normal individuals
of my variety growing between them. The seeds of both were saved and
sown separately, and the two lots of offspring cultivated close to each
other under the same external conditions. In the beginning no difference
was seen, but as soon as the young plants had unfolded three or four
leaves, the progeny of the better nourished half of the parent plant
showed a manifest advance. This difference increased rapidly and was
easily seen in the beds, even before the flowering period.
This experience probably gives an explanation [353] why the
quinquefoliolate variety is so seldom met with in the wild state. For
even if it did occur more often, the plants would hardly find
circumstances favorable enough for the full development of their
varietal character. They must often be so poor in anomalous leaves as to
be overlooked, or to be taken for instances of the commonly occurring
quadrifoliolate leaves and therefore as not indicating the true variety.
Public-domain text, read in full here on John Shaqi.
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