The common cockscomb or _Celosia cristata_, one of the oldest and most
widely cultivated fasciated varieties may be used to illustrate the
first point. In beds it is often to be seen in quite uniform lots of
large and beautiful crests, but this uniformity is only secured by
careful culture and selection of the best individuals. In experimental
trials such selection must be avoided, and in doing so a wide range of
variability at once shows itself. Tall, branched stems with fan-shaped
tops arise, constituting a series of steps towards complete atavism.
This last [412] however, is not to be reached easily. It often requires
several successive generations grown from seed collected from the most
atavistic specimens. And even such selected strains are always reverting
to the crested type. There is no transgression, no springing over into a
purely atavistic form, such as may be supposed to have once been the
ancestor of the present cockscomb. The variety includes crests and
atavists, and may be perpetuated from both. Obviously every gardener
would select the seeds of the brightest crests, but with care the full
crests may be recovered, even from the worst reversionists in two or
three generations. It is a double race of quite the same constitution as
the twisted teasels.
My second point is a direct proof of this assertion, but made with a
fasciated variety of a wild species. I took for my experiment the rough
hawksbeard. In the summer of 1895 I isolated some atavists of the fifth
generation of my race, which, by ordinary selection, gave in the average
from 20-40% of fasciated stems. My isolated atavists bore abundant
fruit, and from these I had the next year a set of some 350 plants, out
of which about 20% had broadened and linear rosettes. This proportion
corresponds with the degree of inheritance which is shown in many years
by the largest and strongest [413] fasciated stems. It strengthens our
conclusion as to the innermost constitution of the double races or
ever-sporting varieties.
Twisted stems and fasciations are very striking monstrosities. But they
are not very good for further investigation. They require too much space
and too much care. The calculation of a single percentage requires the
counting of some hundreds of individuals, taking many square meters for
their cultivation, and this, as my best races are biennial, during two
years. For this reason the countings must always be very limited, and
selection is restrained to the most perfect specimens.
Public-domain text, read in full here on John Shaqi.
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