Leaving these obvious cases of double adaptation, there still remains
one point to be considered. It is the dwarf stature of so many desert
and alpine plants. Are these dwarfs only the extremes of the normal
fluctuating variability, or is their stature to be regarded as the
expression of some peculiar adaptive but latent quality? It is as yet
difficult to decide this question, because statistical studies of this
form of variability are still wanting. The capacity of ripening the seed
on individuals of dwarf stature however, is not at all a universal
accompaniment of a variable height. Hence it cannot be considered as a
necessary consequence of it. On the other hand the dwarf varieties of
numerous garden-plants, as for instance: of larkspurs, snapdragon,
opium-poppies and others are quite stable and thence are obviously due
to peculiar characteristics. Such characteristics, if combined with tall
stature into a pair of antagonists, would yield a double [453]
adaptation, and on such a base a hypothetical explanation could no doubt
be rested. Instead of discussing this problem from the theoretical side,
I prefer to compare those species which are capable of assuming a dwarf
stature under less uncommon conditions than those of alpine and
desert-plants. Many weeds of our gardens and many wild species have this
capacity. They become very tall, with large leaves, richly branched
stems and numerous flowers in moist and rich soil. On bad soil, or if
germinating too late, when the season is drier, they remain very small,
producing only a few leaves and often limiting themselves to one
flower-head. This is often seen with thorn-apples and amaranths, and
even with oats and rye, and is notoriously the case with buckwheat.
Gauchery has observed that the extremes differ often as much from one
another as 1:10. In the case of the Canadian horseweed or _Erigeron
canadensis_, which is widely naturalized in Europe, the tallest
specimens are often twenty-five times as tall as the smallest, the
difference increasing to greater extremes, if besides the main stem, the
length of the numerous branches of the tall plants are taken into
consideration. Other instances studied by the French investigator are
_Erythraea pulchella_ and _Calamintha Acinos_.
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