[454] Dimorphism is of universal occurrence in the whole vegetable
kingdom. In some cases it is typical, and may easily be discerned from
extreme fluctuating variability. In others the contrast is not at all
obvious, and a closer investigation is needed to decide between the two
possibilities. Sometimes the adaptive quality is evident, in other cases
it is not. A large number of plants bear two kinds of leaves linked with
one another by intermediate forms. Often the first leaves of a shoot, or
those of accidentally strong shoots, exhibit deviating shapes, and the
usefulness of such occurrences seems to be quite doubtful. The
elongation of stems and linear leaves, and the reduction of lateral
organs in darkness, is manifestly an adaptation. Many plants have
stolons with double adaptations which enable them to retain their
character of underground stems with bracts or to exchange it for the
characteristics of erect stems with green leaves according to the outer
circumstances. In some shrubs and trees the capacity of a number of buds
to produce either flowers or shoots with leaves seems to be in the same
condition. The capacity of producing spines is also a double adaptation,
active on dry and arid soil and latent in a moist climate or under
cultivation, as with the wild and cultivated apple, and in the
experiments of Lothelier [455] with _Berberis_, _Lycium_ and other
species, which lose their spines in damp air.
In some conifers the evolution of horizontal branches may be modified by
simply turning the buds upside down. Or the lateral branches can be
induced to become erect stems by cutting off the normal summit of a
tree. Numerous organs and functions lie dormant until aroused by
external agencies, and many other cases could be cited, showing the wide
occurrence of double adaptation.
There are, however, two points, which should not be passed over without
some mention. One of them is the influence of sun and shade on leaves,
and the other the atavistic forms, often exhibited during the juvenile
period.
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