Nearly all organs show differences. The most marked are those of the
leaves, which may be small or large, linear or elliptic or oblong and
even rhomboidal in shape, more or less hairy with simple or with
stellate branched hairs, and finally of a pure green or of a glaucous
color. The petals are as a rule obcordate, but this type may be combined
with others having more or less broad emarginations at the summit, and
with differences in breadth which vary from almost linear types to
others which touch along their margins. The pods are short and broad, or
long and narrow, or varying in sundry other [49] ways. All in all there
are constant differences which are so great that it has been possible to
distinguish and to describe large numbers of types.
Many of them have been tested as to their constancy from seed. Jordan
made numerous cultures, some of which lasted ten or twelve years; Thuret
has verified the assertion concerning their constancy by cultures
extending over seven years in some instances; Villars and de Bary made
numerous trials of shorter duration. All agree as to the main points.
The local races are uniform and come true from seed; the variability of
the species is not of a fluctuating, but of a polymorphous nature. A
given elementary species keeps within its limits and cannot vary beyond
them, but the whole group gives the impression of variability by its
wide range of distinct, but nearly allied forms.
The geographic distribution of these elementary species of the
whitlow-grass is quite distinct from that of the violets. Here
predominant species are limited to restricted localities. Most of them
occupy one or more departments of France, and in Holland two of them are
spread over several provinces. An important number are native in the
centre of Europe, and from the vicinity of Lyons, Jordan succeeded in
establishing about fifty elementary [50] species in his garden. In this
region they are crowded together and not rarely two or even more quite
distinct forms are observed to grow side by side on the same spot.
Farther away from this center they are more widely dispersed, each
holding its own in its habitat. In all, Jordan has distinguished about
two hundred species of _Draba verna_ from Europe and western Asia.
Subsequent authors have added new types to the already existing number
from time to time.
The constancy of these elementary species is directly proven by the
experiments quoted above, and moreover it may be deduced from the
uniformity of each type within its own domain. These are so large that
most of the localities are practically isolated from one another, and
must have been so for centuries. If the types were slowly changing such
localities would often, though of course not always, exhibit slighter
differences, and on the geographic limits of neighboring species
intermediates would be found. Such however, are not on record. Hence the
elementary species must be regarded as old and constant types.
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