One of them is the hawkweed or _Hieracium_, and the other is the
dandelion or _Taraxacum officinale_. Hawkweeds are known as a genus in
which the delimitation of the species is almost impossible, Thousands of
forms may be cultivated side by side in botanical gardens, exhibiting
[60] slight but undoubted differentiating features, and reproduce
themselves truly by seed. Descriptions were formerly difficult and so
complicated that the ablest writers on this genus, Fries and Nageli are
said not to have been able to recognize the separate species by the
descriptions given by each other. Are these types to be considered as
elementary species, or only as individual differences? The decision of
course, would depend upon their behavior in cultures. Such tests have
been made by various experimenters. In the dandelion the bracts of the
involucre give the best characters. The inner ones may be linear or
linear-lanceolate, with or without appendages below the tip; the outer
ones may be similar and only shorter, or noticeably larger, erect,
spreading or even reflexed, and the color of the involucre may be a pure
green or glaucous; the leaves may be nearly entire or pinnatifid, or
sinuate-dentate, or very deeply runcinate-pinnatifid, or even pinnately
divided, the whole plant being more or less glabrous.
Raunkiaer, who has studied experimentally a dozen types from Denmark,
found them constant, but observed that some of them have no pollen at
all, while in others the pollen, though present, is impotent. It does
not germinate on the stigma, cannot produce the ordinary tube, [61] and
hence has no fertilizing power. But the young ovaries do not need such
fertilization. They are sufficient unto themselves. One may cut off all
the flowers of a head before the opening of the anthers, and leave the
ovaries untouched, and the head will ripen its seeds quite as well. The
same thing occurs in the hawkweeds. Here, therefore, we have no
fertilization and the extensive widening of the variability, which
generally accompanies this process is, of course, wanting. Only partial
or vegetative variability is present. Unfertilized eggs when developing
into embryos are equivalent to buds, separated from the parent-plant and
planted for themselves. They repeat both the specific and the individual
characters of the parent. In the case of the hawkweed and the dandelion
there is at present no means of distinguishing between these two
contrasting causes of variability. But like the garden varieties which
are always propagated in the vegetative way, their constancy and
uniformity are only apparent and afford no real indication of hereditary
qualities.
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