Granting this state of things as the only probable one, we can easily
imagine what must have been the consequences. If a wild species had been
taken into cultivation only once, the cultivated form would have been a
single elementary [65] type. But it is not very likely that such
partiality would occur often. The conception that different tribes at
different times and in distant countries would have used the wild plants
of their native regions seems far more natural than that all should have
obtained plants for cultivation from the same source or locality. If
this theory may be relied upon, the origin of many of the more widely
cultivated agricultural plants must have been multiple, and the number
of the original elementary species of the cultivated types must have
been so much the larger, the more widely distributed and variable the
plants under consideration were before the first period of cultivation.
Further it would seem only natural to explain the wide variability of
many of our larger agricultural and horticultural stocks by such an
incipient multiformity of the species themselves. Through commercial
intercourse the various types might have become mixed so as to make it
quite impossible to point out the native localities for each of them.
Unfortunately historical evidence on this point is almost wholly
lacking. The differences in question could not have been appreciated at
that remote period, and interest the common observer but little even
today. The history of most of the cultivated plants is very obscure,
[66] and even the most skillful historians, by sifting the evidence
afforded by the older writers, and that obtained by comparative
linguistic investigations have been able to do little more than frame
the most general outline of the cultural history of the most common and
most widely used plants.
Some authors assume that cultivation itself might have been the
principal cause of variability, but it is not proved, nor even probable,
that cultivated plants are intrinsically more variable than their wild
prototypes. Appearances in this case are very deceptive. Of course
widely distributed plants are as a rule richer in subspecies than forms
with limited distribution, and the former must have had a better chance
to be taken into cultivation than the latter. In many cases, especially
with the more recent cultivated species, man has deliberately chosen
variable forms, because of their greater promise. Thirdly, wide
variability is the most efficient means of acclimatization, and only
species with many elementary units would have offered the adequate
material for introduction into new countries.
From this discussion it would seem that it is more reasonable to assert
that variability is one of the causes of the success of cultivation,
than to assume that cultivation is a cause of variability [67] at large.
And this assumption would be equally sufficient to explain the existing
conditions among cultivated plants.
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