The fleshiness or pulpiness of the roots is still more variable. Some
are as thick as the arm and edible, others are not thicker than a finger
and of a woody composition, and the structure of this woody variety is
very interesting. The sugar-beet consists, as is generally known, of
concentric layers of sugar-tissue and of vascular [69] strands; the
larger the first and the smaller the latter, the greater is, as a rule,
the average amount of sugar of the race. Through the kindness of the
late Mr. Rimpau, a well known German breeder of sugar-beet varieties, I
obtained specimens from seed of a native wild locality near Bukharest.
The plants produced quite woody roots, showing almost no sugar tissue at
all. Woody layers of strongly developed fibrovascular strands were seen
to be separated one from another only by very thin layers of
parenchymatous cells. Even the number of layers is variable; it was
observed to be five in my plants; but in larger roots double this number
and even more may easily be met with.
Some authors have distinguished specific types among these wild forms.
While the cultivated beets are collected under the head of _Beta
vulgaris_, separate types with more or less woody roots have been
described as _Beta maritima_ and _Beta patula_. These show differences
in the habit of the stems and the foliage. Some have a strong tendency
to become annual, others to become biennial. The first of course do not
store a large quantity of food in their roots, and remain thin, even at
the time of flowering. The biennial types occur in all sizes of roots.
In the annuals the stems may vary from [70] erect to ascending, and the
name _patula_ indicates stems which are densely branching from the base
with widely spreading branches throughout. Mr. Em. von Proskowetz of
Kwassitz, Austria, kindly sent me seeds of this _Beta patula_, the
variability of which was so great in my cultures as to range from nearly
typical sugar-beets to the thin woody type of Bukharest.
Broad and narrow leaves are considered to be differentiating marks
between _Beta vulgaris_ and _Beta patula_, but even here a wide range of
forms seem to occur.
Rimpau, Proskowetz, Schindler and others have made cultures of beets
from wild localities in order to discover a hypothetical common ancestor
of all the present cultivated types. These researches point to the _B.
patula_ as the probable ancestor, but of course they were not made to
decide the question as to whether the origination of the several now
existing types had taken place before or during culture. From a general
point of view the variability of the wild species is parallel to that of
the cultivated forms to such a degree as to suggest the multiple origin
of the former. But a close investigation of this highly important
problem has still to be made.
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