Leaving aside all these different types, we are concerned here only with
the "seed-varieties" of pure origin, or at least with those, that are
supposed to be so. Hybridization and vegetative multiplication of the
hybrids no doubt occur in nature, but they are very rare, when compared
with the ordinary method of propagation by seed. "Seed-varieties" may
further be divided into constant and inconstant ones. The difference is
very essential, but the test is not always easy to apply. Constant
varieties are as sharply defined and as narrowly limited as are the best
wild species, while inconstant types are cultivated chiefly on account
of their wide range of form and color. This diversity is repeated
yearly, even from the purest seed. We will now discuss the constant
seed-varieties, leaving the inconstant and eversporting types to a
subsequent lecture.
In this way we may make an exact inquiry into the departures from the
species which are ordinarily considered to constitute the essential
character of such a constant and pure seed-variety [124] and need only
compare these differences with those that distinguish the elementary
species of one and the same group from each other.
Two points are very striking. By far the greatest part of the ordinary
garden-varieties differ from their species by a single sharp character
only. In derivative cases two, three or even more such characters may be
combined in one variety, for instance, a dwarfed variety of the larkspur
may at the same time bear white flowers, or even double white flowers,
but the individuality of the single characters is not in the least
obscured by such combinations.
The second point is the almost general occurrence of the same variety in
extended series of species. White and double flowers, variegated leaves,
dwarfs and many other instances may be cited. It is precisely this
universal repetition of the same character that strikes us as the
essential feature of a variety.
And again these two characteristics may now be considered separately.
Let us begin with the sharpness of the varietal characters. In this
respect varieties differ most obviously from elementary species. These
are distinguished from their nearest allies in almost all organs. There
is no prominent distinctive feature between the single forms of _Draba_
[125] _Verna_, _Helianthemum_ or of _Taraxacum_; all characters are
almost equally concerned. The elementary species of _Draba_ are
characterized, as we have seen, by the forms and the hairiness of the
leaves, the number and height of the flower-stalks, the breadth and
incision of the petals, the forms of the fruits, and so on. Every one of
the two hundred forms included in this collective species has its own
type, which it is impossible to express by a single term. Their names
are chosen arbitrarily. Quite the contrary is the case with most of the
varieties, for which one word ordinarily suffices to express the whole
difference.
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