To a sub-class selected for a centre, two qualities are indispensably
requisite. They have been previously given from Mr. MacLeay, who, it
will be remembered, discovered that one of each of his five groups
contained types of the other four, besides a type peculiar to itself.
This quality must hold good in any group thus selected for a centre;
it must contain types of the six surrounding groups in the first
place. Now, is this applicable to Papilio? Have we not already
experienced the greatest difficulty in finding three good approaches,
the smallest number which a sub-class can possess? How then can we
hope, by any good fortune in discovery, to make ourselves masters of
three other entirely new ones, and these to sub-classes to which it is
confessedly the most unlike? Phalæna, on the contrary, presents us
with Lasiocampa, Ægeria, Cilix, Lithosia, Apatela and Orgyia, five of
which genera beautifully typify the approximating sub-classes. The
preference on this score then is decidedly with Phalæna.
The second position, that it should contain a type peculiar to itself,
is almost a matter of course; but my own idea is, that the very centre
should not only be a type of the genus, or order, or sub-class, but of
the class itself of which it is the centre. From this position, then,
a further and still more important question arises,--What is the type
of Lepidoptera? The parts which afford the generic characters of
Lepidoptera, and, I believe, generic characters in the perfect state
are the only ones of any value, are these--the mouth, palpi, antennæ
and wings; and, as no medium can constitute a type, the excess of
these characters, whether superlatively or diminutively considered,
must be resorted to as the most probable means we possess of
discovering what this type may really be. First, then, the mouth. In
Lepidoptera, we find two distinct characters in this;--first, its
entire absence; secondly, its being furnished with prodigiously long
antlia. The first character is that of Phalæna, the second that of
Sphinx. Next, the palpi are either entirely obsolete or exceedingly
prominent, the first in Phalæna, the second in Pyralis. Thirdly, the
antennæ are remarkably pectinated, or clavated, or setaceous: the
first character is that of Phalæna, the second that of Papilio, the
third that of Noctua. Fourthly, the wings are enormously expansive in
proportion to the body, or remarkably small,--the first is the
character of Phalæna, the second that of Sphinx. It need scarcely be
added, that all these characters are to be met with in every
intermediate degree of intensity. Now, it appears, that Phalæna
possesses an extreme of each of the four principal characters, Sphinx
of two, Noctua of one, and Papilio of one; therefore Phalæna is the
typical genus, Phalænæ the typical order, and Phalæna the typical
sub-class of Lepidoptera: and a necessary conclusion from this fact
is, the type of Lepidoptera is an insect without antlia or palpi, with
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