The berries after drying are spherical and about one-fifth of an inch in
diameter and are wrinkled on the surface, indistinctly pointed below by
the remains of a very short pedical and crowned by three or four lobed
stigmas. The thin pericarp tightly encloses a single seed, the embryo of
which, on account of the premature gathering is not fully developed and
is replaced by the cavity below the apex. The seed itself contains
within the thin red-brown testa a shining albumen of angular, radically
arranged, large-celled parenchyme, gray and horny without and mealy
within.
The transverse section of a grain of pepper exhibits a soft, yellowish
epidermis covering; the outer pericarp is formed of a closely packed
yellow layer of large and most radically arranged thick-walled cells,
most of which are colorless and loaded with starch; others contain a
soft, yellowish, amorphous mass, each containing in its minute cavity a
quantity of dark-brown resin, while the middle layer of the pericarp
consists of starch and oil, the shrinkage of which causes the deep
wrinkles on the surface of the berry. The next inner layer of the
pericarp exhibits its circumference tangentially arranged soft
parenchyme, the cells of which possess either spiral striation or spiral
fibers, but towards the interior lose parenchyme free from starch and
containing very large oil cells. The testa is formed in the first place
of a row of small yellowish thick-walled cells, next to which follows
the true testa as a dense, dark-brown layer of lignified cells, the
individual outlines of which are indistinguishable. If thin slices are
kept under glycerine for some time these masses are slowly transformed
into needle-shaped crystals of piperine. The angular cells of the
interior of the seed are, of course, the more prominent and, when once
seen, their characteristic form and contents are easily recognized
again. The structure of the outer coats is made out with more
difficulty, and before attempting to do so on ground pepper it is best
to soften some whole black and white pepper corns in glycerine and cut
sections from various parts of the exterior of the berry.
Public-domain text, read in full here on John Shaqi.
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