We have seen that the class Crustacea (crabs, shrimps, etc.) is the
great division of the Arthropoda entirely adapted to an aquatic life,
breathing, by means of gills, the air which is dissolved in the water.
Insects and spiders are air-breathing, and properly belong to the land;
yet there are many insects which pass their early stages—often the
greater portion of their life—in the water, and some which are very
fairly at home there when adult. Such insects often have gills when
young, and are therefore at that period true water animals, like the
Crustacea.
The Arachnida—that division of the Arthropoda to which the spiders
belong—include a few groups which permanently inhabit the sea, and
could not live on land. There are even some weird creatures called
Sea-spiders (Pycnogonids), but these do not concern us, for they are
very far removed from the true spiders which are the subject of our
investigations.
Now the true spiders are always air-breathing, and if they venture
into the water at all they must frequently come up to the surface
to breathe, or else they must store up a reservoir of air beneath
the surface of the water if they are to avoid death by drowning.
Nevertheless some of them have been hardy enough to encroach on the
domain of the Crustacea. Not a few are able to run freely on the
surface of the water and even to dive occasionally for the purpose
of seizing one of its denizens, but the number of those which have
succeeded in really adapting themselves to aquatic life is very
limited, and is, as far as we know, restricted to two small groups,
both of them members of the Agelenidae.
Public-domain text, read in full here on John Shaqi.
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