The other type of trap-door is less interesting and much more
elementary, consisting simply of a wafer-like sheet of silk mixed with
earth and vegetable matter, but it is a curious fact that while all
known trap-door nests of the cork type are simple tubes, the burrows
with wafer doors are often much more complex. In some cases there
is a branch tube, like that constructed by _Lycosa picta_, leaving
the main tunnel at a depth of some three inches, and reaching the
surface perhaps two inches away from the trap-door, so that the whole
excavation is +Y+-shaped. This branch tube is permanently closed
by a thin sheet of silk and earth, which, however, it would not be
difficult to break through if it were urgent for the spider to escape
while the enemy was exploring the main tunnel.
But a more interesting case is the occurrence of another trap-door
some way down the tube. If the tube is unbranched, this forms merely a
second line of defence if the outer door is forced, but in the case of
a branched tube the additional door hangs at the fork of the +Y+,
and is so shaped as to form a perfect valve, so that the spider, by
holding it against one or the other side of the tunnel, can connect the
bottom limb of the +Y+ with either fork at will, leaving to the
intruder a beautifully smooth-lined tube to explore, with no hint of
the possibility of escape in other directions.
There are sometimes other complications in the ramification of the
tube, but these need not detain us. Each species of spider adheres to
its own particular type of architecture, and may safely—in a given
neighbourhood—be identified by its nest.
As with the Lycosidae, the burrowing is all done by the mandibles,
but here the first joint—the handle of the penknife—is of more
importance than the blade or fang. Indeed the burrowing species of the
Aviculariidae may be distinguished from the rest by their mandibles,
which are provided in front with a _rastellum_, or row of teeth for
digging. A trap-door spider, then, does not go to work like a rabbit,
or a terrier, scratching and kicking away the earth as it digs; it
laboriously dislodges particles of soil with its powerful mandibles,
and carries away the loosened fragments to deposit them at a distance.
The trap-door spiders of the Mediterranean region are nocturnal
creatures, and little is known of their habits. Erber relates that a
species found in the island of Tinos comes out at night, fixes open the
trap-door with a few threads, and spins a web near its nest to entrap
passing insects, clearing away any trace of it before the dawn. In the
case of some Chinese and also some Australian species observers allege
that they frequently wander from their nests in the day-time.
Public-domain text, read in full here on John Shaqi.
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