The spinnerets, then, are only the bearers of the actual tubes which
emit the silk. The distribution of the tubes themselves is different
in the different kinds of spiders, but it is usually possible to
distinguish two kinds. There are generally present a large number of
very fine cylindrical tubes or “spools” and a few conical tubes of much
larger base, which are called spigots. Each of these orifices, whether
on spool or spigot, is connected by a fine tube with a separate silk
gland, or organ for manufacturing silk, situated within the spider’s
abdomen. Epeira has about 600 of such glands, each with its own
terminal spool or spigot, and the large number of these tubes has given
rise to a misconception that is very widely spread—namely that the
spider’s line, fine as it is, is “woven” of hundreds of threads of very
much finer silk. This is not so, as we shall presently see.
Though Epeira has some 600 silk-glands, it has only five different
_kinds_ of gland, manufacturing silk of different properties. No other
family of spiders has so many, though two other kinds of gland have
been found in less elaborate spinners. Within the spider the silk is
fluid but it solidifies on meeting the air, each thread hardening as it
emerges though still continuous with the fluid contents of the gland,
so that the drawing out of a silken thread is just like the operation
so familiar with the glue-pot, or with spun glass, except that the
hardening is not due to cooling but to exposure to the air. This
general description will, it is hoped, make an account of the organs in
Epeira more comprehensible.
The spinnerets of Epeira are so small and inconspicuous that their
disposition is not very easy to make out. When not in use they form
a tiny cone under the tip of the abdomen, and only four are visible,
their free ends being so brought together as entirely to conceal a
small central pair. There are really, then, three pairs of spinnerets
which we may call at once the anterior, median and posterior pairs,
though when at rest only the anteriors and posteriors can be seen.
If the spider is observed with a pocket-lens as it crawls about in a
glass tube it will be noticed that the spinnerets are capable of great
mobility. Their ends can be separated or brought together, or they may
be made to rub against each other or against the sides of the tube.
The anteriors and posteriors, moreover, are two-jointed though the
medians consist only of a single joint.
So much can be seen without any great magnification, but the microscope
will be necessary if a complete understanding of their mechanism is to
be arrived at. What it reveals will now be briefly described, and will,
it is hoped, be made tolerably clear by the accompanying figures which
are simplified by the omission of a large number of bristles which tend
to hide the essential structure, and by a great reduction in the number
of “spools,” though the spigots are all indicated.
Public-domain text, read in full here on John Shaqi.
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