GRADE IV. _The Coelomata or Triploblastic Animals with a
Body-Cavity._--This grade includes all the remainder of the animal
kingdom. As an example of it, we may take the Common Frog. If we open
from the lower surface the dead body of a frog, we first cut through
the skin, next the muscles; then we come to the viscera, lying neatly
packed in a cavity from which we can dislodge them. This cavity is the
Body-Cavity. The skin corresponds with the ectoderm of Hydra, although
it is a vastly more complicated affair. The glandular lining of the
alimentary canal corresponds with the endoderm of Hydra; although this,
too, is a more complicated affair. The mass of the body, lying between
these two layers, is considered to correspond somewhat with the mesoderm
of Grade III., and has received the collective term of Mesoblast. This
description applies equally to the earthworm, for the higher worms
differ immensely from the lower worms, and stand on a level with more
important members of the animal kingdom (see Fig. 41, p. 139). The
body-cavity may be formed in different ways in different animal groups;
but there is reason to believe that in certain cases it originates by a
folding off of part of an original cavity corresponding with that of
Hydra; so that part went to form the intestine, and part the cavity
surrounding it.
The above arrangement of the main great groups of animals into four
grades is that given by Professor Arnold Lang.
It should be added, that there are a few exceptional forms that present
a departure from these broad rules of structure. They are, however, so
few that they need only be named as curiosities. For instance, there are
parasites in which the inner body-layer is practically done away with,
because they are fitted to absorb food through the outer layer. And in
one division of the Moss-Corals there is no body-cavity to be seen,
although it is to be found in the other division.
What is the outcome of all this sorting of the animal kingdom? This most
important result: that a classification of the animal kingdom into the
four grades we have named, presents, in serial order, the stages through
which young animals of the higher forms pass in the course of their
growth. Every creature begins as a unicellular organism--the fertilised
egg-cell. A vast number of creatures belonging to the higher groups
present, later on, a two-layered condition, comparable with that of
Grade II. Later on they acquire a third layer, and therefore correspond
with Grade III. By degrees the body-cavity is formed, and they then
present the adult body-structure of Grade IV. The development of the
chicken in the egg, for instance, presents these four stages.
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