Strange to say, one of these structures is the ear. This takes its
origin from one of the gill-"clefts" or spaces. The Eustachian tube,
which communicates between the ear and the nose, is part of this cleft;
and the little bones which are inside the ear represent the bones
of that gill-cleft. For, in fishes, bones support each gill, and
are connected together to form a complex arrangement. In the higher
vertebrates, which possess gills only in the embryo, this gill-skeleton
is much modified, and persists as a bone, the hyoid bone supporting the
tongue.
The gills of vertebrates, arranged in successive pairs along the throat,
are "perforating gills"; that is to say, they consist essentially of
holes or spaces which pass right through the wall of the throat.
If we were to seek for a general character of the vertebrates, besides
those mentioned above, that they all possess a notochord and gills, we
might also find it in the character of the skin. Fishes, Reptiles, Birds
and Mammals, all agree in this, that they have a special clothing of the
skin--scales, feathers and fur, respectively. These three kinds of
structure, although so widely differing in appearance, are practically
formed all in the same way, viz., by alternate ingrowths and outgrowths
of the skin; the ingrowth forming the root of the scale, hair or
feather, and the outgrowth its projecting part. If these infoldings and
outgrowths of the skin could be straightened out into a plane surface,
the skin of a small vertebrate would cover an enormous area. The above
list excludes the Amphibia: in this class, it should be mentioned, the
scales have been lost, and are only found in one group.
The scales of Fishes were at one time proposed as a basis of
classification: large groups being characterized respectively by the
possession of plain rounded scales (cycloids), scales fringed at the
posterior end (ctenoid, or comb-like); placoid scales, consisting of
bony plates, and ganoid scales, large plates covered with shiny enamel.
These distinctions, however, were not found useful as a guide in
classification. The diagram shows the elaborate scales of the common
sole.
[Illustration: FIG. 42.--Scales of the Common Sole, highly magnified.]
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