Let us now consider some other creatures that resemble vertebrates in
some ways, and help to form the group of Chordata. Balanoglossus is one
of them, the Acorn-tongue Animal. This odd name is given to it on
account of a structure which is called (like the elephant's trunk) a
Proboscis; this may be compared with a tongue, so far as its use goes,
for it is thrust out to catch prey and again drawn in. It is oval in
shape, and therefore fancifully compared to an acorn. It is highly
sensitive, being richly supplied with nerves. The creature is to all
intents and purposes a kind of worm; and, like many of the higher
worms, it has a larva with bands of cilia. This larva, which is better
represented in some species than in others, was originally described
under the name of _Tornaeria_. It is considered to resemble, in some
degree, the larva of Echinoderms; on this hint, some zoologists have
sought to establish a connection between Vertebrates and Echinoderms,
and have been able to find other points of comparison besides the one
named. It remains to be seen whether this suggestion will lead to
further results. It may be added that the larva of Balanoglossus has
also been compared with that of Phoronis (p. 122), thus assuming a
relationship with the Polyzoa, and through them with the Brachiopoda. It
appears, therefore, that the subject of the possible relationships of
the Vertebrata is one of the greatest complexity. The last named theory,
however, has been adversely criticised by very high authority.
We have not, however, explained yet what is the claim of Balanoglossus
to be grouped with the Chordata. This consists in the fact that a
certain part associated with the interior of the proboscis has been
identified, from its structure, mode of origin, and relations with
the nerves, as a notochord. Balanoglossus also agrees with the true
vertebrates in possessing successive pairs of perforating gills (see p.
142), which are especially noticeable in the young animal. The presence
of this feature is important, in view of the fact that some authorities
have sought to throw doubt on the genuineness of the notochord of
Balanoglossus.
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