Experiment thus confirmed the view that _Bryoniæ_ is the parent-form
of _Napi_, and the description hitherto given by systematists ought
therefore properly to be reversed. _Pieris Bryoniæ_ should be elevated
to the rank of a species, and the ordinary winter and summer forms
should be designated as vars. _Napi_ and _Napeæ_. Still I should not
like to take it upon myself to increase the endless confusion in the
synonomy of butterflies. In a certain sense, it is also quite correct
to describe the form _Bryoniæ_ as a climatic variety, for it is, in
fact, established, if not produced, by climate, by which agency it
is likewise preserved; only it is not a secondary, but the primary,
climatic variety of _Napi_. In this sense most species might probably
be described as climatic varieties, inasmuch as under the influence of
another climate they would gradually acquire new characters, whilst,
under the influence of the climate now prevailing in their habitats,
they have, to a certain extent, acquired and preserved their present
form.
The var. _Bryoniæ_ is, however, of quite special interest, since it
makes clear the relation which exists between climatic variation
and seasonal dimorphism, as will be proved in the next section. The
correctness of the present theory must first here be submitted to
further proof.
It has been shown that the secondary forms of seasonally dimorphic
butterflies do not all possess the tendency to revert in the same
degree, but that this tendency rather varies with each individual. As
the return to the primary form is synonymous with the relinquishing of
the secondary, the greater tendency to revert is thus synonymous with
the greater tendency to relinquish the secondary form, but this again
is equivalent to a lesser stability of the latter; it must consequently
be concluded that the individuals of a species are very differently
influenced by climatic change, so that with some the new form must
become sooner established than with others. From this a variability of
the generation concerned must necessarily ensue, i.e., the individuals
of the summer generation must differ more in colour and marking than
is the case with those of the winter generation. If the theory is
correct, the summer generations should be more variable than the winter
generations--at least, so long as the greatest possible equalization of
individual variations has not occurred through the continued action of
warmth, combined with the constant crossing of individuals which have
become changed in different degrees. Here also the theory is fully in
accord with facts.
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