If, as I have attempted to show, seasonal dimorphism originates
through the slow operation of a changed summer climate, then is this
phenomenon nothing else than the splitting up of a species into two
climatic varieties in the same district, and we may expect to find
various connexions between ordinary simple climatic variation and
seasonal dimorphism. Cases indeed occur in which seasonal dimorphism
and climatic variation pass into each other, and are interwoven in
such a manner that the insight into the origin and nature of seasonal
dimorphism gained experimentally finds confirmation. Before I go more
closely into this subject, however, it is necessary to come to an
understanding as to the conception “climatic variation,” for this term
is often very arbitrarily applied to quite dissimilar phenomena.
According to my view there should be a sharp distinction made between
climatic and local varieties. The former should comprehend only such
cases as originate through the direct action of climatic influences;
while under the general designation of “local forms,” should be
comprised all variations which have their origin in other causes--such,
for example, as in the indirect action of the external conditions of
life, or in circumstances which do not owe their present existence to
climate and external conditions, but rather to those geological changes
which produce isolation. Thus, for instance, ancient species elsewhere
long extinct might be preserved in certain parts of the earth by the
protecting influence of isolation, whilst others which immigrated in
a state of variability might become transformed into local varieties
in such regions through the action of ‘amixia,’[24] i.e. by not being
allowed to cross with their companion forms existing in the other
portions of their habitat. In single cases it may be difficult, or for
the present impossible, to decide whether we have before us a climatic
form, or a local form arising from other causes; but for this very
reason we should be cautious in defining climatic variation.
The statement that climatic forms, in the true sense of the word, do
exist is well known to me, and has been made unhesitatingly by all
zoologists; indeed, a number of authentically observed facts might
be produced, which prove that quite constant changes in a species may
be brought about by the direct action of changed climatic conditions.
With butterflies it is in many cases possible to separate pure climatic
varieties from other local forms, inasmuch as we are dealing with
only unimportant changes and not with those of biological value, so
that natural selection may at the outset be excluded as the cause of
the changes in question. Then again the sharply defined geographical
distribution climatically governed, often furnishes evidence of
transition forms in districts lying between two climatic extremes.
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